Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
Prof. Zeiller[1010] described this species as _Cardiocarpus indicus_
from the Karharbari (Lower Gondwana) beds of India. An examination of
the type-specimens enables me to confirm the accuracy of the original
account. The platyspermic seed is 5·5 cm. long and 4·5 cm. broad;
a cordate nucule is enclosed by a flat border similar to that of
_Samaropsis Leslii_ but narrower especially on the sides of the nucule.
At the apex there is a deep sinus extending to the nucule, and at the
base a fairly broad band of carbonaceous matter shows the position of
the chalazal vascular strand. The seed is characterised by its large
size and by its almost orbicular form: it occurs as a detached specimen
in beds containing _Cordaites_ (_Noeggerathiopsis_) and _Glossopteris_.
A species from Arkansas described by Lesquereux[1011] as _Cardiocarpus
ingens_ [= _Cordaicarpus ingens_ (Lesq.)] affords another example of a
large seed similar to _C. indicus_ and, except in its more orbicular
form, to _Samaropsis Leslii_.
_Samaropsis Milleri_ (Feistmantel).
[Illustration: Fig. 504. _Samaropsis Milleri._ (Indian Geological
Survey, Calcutta; nat. size.)]
An examination of the type-specimen from the Calcutta Museum enables
me to amplify the original description in an important particular. The
species was found in Lower Gondwana (Katharbari) beds in India and
referred by Feistmantel[1012] to the genus _Carpolithes_. Arber[1013],
who tentatively employed the generic name _Cardiocarpus_ in place
of _Carpolithes_, suggests that the seeds may be radiospermic, as
Feistmantel’s drawings show only a very narrow border to the nucule.
The specimen reproduced in fig. 504 was figured by Feistmantel without
any indication of a definite sarcotesta or wing, but as seen in the
drawing the ovate cordate sclerotesta is surrounded on one side and at
the base by an outer envelope: this is clearly seen at the apex where
it shows a rounded termination sloping downwards towards the micropyle
precisely as in _Samaropsis indica_ (Zeill.). The border is narrow at
the sides and broader at the base as in _S. Leslii_. The seed is 4·5
cm. long and 2·3 cm. broad, differing from _S. indica_ in its rather
smaller size and in the slightly narrower nucule. Though there is no
decisive evidence as to the parent-plant the occurrence of a specimen
of this species partially covered by a scale-leaf of a type[1014]
very similar to that which is generally recognised as belonging to
_Glossopteris_ suggests the possibility that the seeds may belong to
that genus. Scale-leaves of _Glossopteris_ are described in Volume
+ii.+, but it may be added here that leaves similar in form to those
from India, Australia, and elsewhere are figured by Geinitz[1015]
from the Altai Mountains as _Trigonocarpus_? _actaeonelloides_: the
specimens are represented with the basal scar at the apex.
=Cordaicarpus.= Geinitz.
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