Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
In view of the fact that the generic names _Cardiocarpus_,
_Cordaicarpus_, and other designations have been applied to casts
and impressions which cannot be distinguished by any constant or
important feature it is proposed to adopt the name _Cordaicarpus_
for platyspermic seeds, preserved as casts or impressions, having
a comparatively narrow border enclosing an ovate or cordate-ovate
nucule; the base is either rounded or cordate. The choice between
_Cordaicarpus_ and _Samaropsis_ depends on the breadth of the border.
_Cordaicarpus_, though more suggestive of a Cordaitean alliance, may in
some cases be a Pteridosperm seed.
_Cordaicarpus Cordai_ (Geinitz). Lenticular seeds more or less
orbicular or broadly ovate (fig. 502, C, D), often slightly cordate at
the base of the nucule and with a broadly acute apex: the border is
narrow or sometimes hardly represented as in the seeds described by
Berger as _Rhabdocarpus ovoides_, a species similar to but more oval
than _Cordaicarpus Cordai_. The latter species[1016] occurs in several
coalfields in Britain, France, Germany, and elsewhere. Fig. 502, C,
shows a good example from the Middle Coal Measures of Yorkshire, 9 mm.
long by 8 mm. broad; on the very slightly cordate nucule are several
faint ribs converging towards the base and apex and between them
fine striations, characters too indistinct to be reproduced in the
natural-size drawing. The flat border represents the sclerotesta. The
seed shown in fig. 502, D, from the Westphalian series of Warwickshire
belongs to the same species or is a closely allied type: the faint
suggestion of reticulation on its surface might be regarded as a reason
for referring it to _C. areolatus_ Boul.[1017], a form characterised
by a reticulation on the testa, described by Zeiller[1018] and other
authors. This reticulation is, however, in some cases at least, formed
by crumpling and splitting of the superficial carbonised film into more
or less regular meshes: the figured specimen occurs with several other
seeds of the same type, most of which have a smooth surface. Dr Kidston
tells me that a recent critical examination of seeds in his collection
leads him to regard some specimens (_e.g._ fig. 502, D) previously
referred by him to _C. Cordai_ as identical with _Carpolithes
membranaceus_ Goepp.[1019]
[Illustration: Fig. 505. _Cordaicarpus Cordai._ Cuticle of the testa.
(Kidston Collection.)]
The photograph reproduced in fig. 505 shows the result of chemically
treating the carbonised cuticle of a seed of _Cordaicarpus Cordai_, a
method little used as yet in the case of Palaeozoic plants but which
may be useful in distinguishing seeds which cannot be satisfactorily
separated by microscopic features. The superficial cells have very
thick walls and present an appearance similar to that of the sclerous
cells in the testas of some petrified specimens.
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