Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The term Cycadophyta, suggested by Nathorst[1069], is used in a
comprehensive sense to include both recent Cycads and the much
larger number of extinct types which it is customary to speak of
as Cycadean plants or fossil Cycads. The designation ‘Cycads’ in
the case of the majority of the fossil forms is, however, open to
criticism on the ground that they differ too widely from existing
genera to be associated with them in one class. It would be pedantic
and inconvenient to give up the almost universal practice of extending
the term Cycad beyond the limits defined by the characters of
recent species. The important point is to adopt some classification
which gives expression to our views as to the degree of affinity
between recent and extinct types. In a considerable number of cases,
especially impressions of presumably Cycadean fronds and stems that
occur without any fertile shoots, it is impossible to determine the
degree of relationship to modern types. It was with a view to a
rational group-designation for such fossils that Nathorst proposed
the term Cycadophyta, including in it the two classes Cycadales and
Bennettitales. The Cycadales comprise the recent genera and such
extinct types as may legitimately be included in the same class, but
as we shall see later there are hardly any fossil species that can be
assigned to this section on thoroughly satisfactory grounds. The term
Bennettitales is used by Engler, Nathorst, and several other authors as
a class-designation for a large number of Mesozoic Cycads agreeing in
their more important morphological characters with the Lower Cretaceous
stems on which Carruthers[1070] founded the genus _Bennettites_,
placing it in a new tribe, the Bennettiteae. Some authors have followed
Carruthers in the use of the family-name Bennettiteae for a subdivision
of the Cycadales equal in rank to the Cycadaceae, while others, wishing
to give greater emphasis to the difference between the extinct and
recent plants, prefer to adopt the class-name Bennettitales. Carruthers
regarded _Bennettites_ as occupying the same position in relation to
other Cycads as _Taxus_ holds with regard to the cone-bearing members
of the Coniferae. The adoption of Bennettitales is intended to convey
the impression that the class is more distantly related to the recent
Cycads than is implied by the analogy of _Taxus_. Nathorst[1071], as
the result of his discovery of certain reproductive organs associated
with some Rhaetic fronds described as _Dioonites spectabilis_, proposed
a third subdivision, the Dioonitales, but he subsequently[1072]
restored the fronds to their original designation _Nilssonia
pterophylloides_, and on further examination found that the supposed
microspore-bearing organs were seeds. _Nilssonia pterophylloides_
would therefore find a more natural place in the class Nilssoniales
instituted by Thomas[1073].
BENNETTITALES.
=Cycadeoidea.= Buckland.
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