Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
In 1827 Buckland[1074] proposed the generic name _Cycadeoidea_ for
some petrified stems from the Purbeck beds in the Isle of Portland and
published a description of two species, _Cycadeoidea megalophylla_
and _C. microphylla_. Brongniart[1075] considered Buckland’s term
_Cycadeoidea_ inappropriate and proposed in its place _Mantellia_, the
type-species being _Mantellia nidiformis_ Brongn., the Portland stem
which Buckland a month or two later called independently _Cycadeoidea
megalophylla_. Subsequently Brongniart[1076] withdrew _Mantellia_ as
it had been previously used by Parkinson for a sponge and substituted
_Cycadites_. Some years later Carruthers[1077] revived _Mantellia_
for a type of Cycadean stem from Portland though one species,
apparently indistinguishable from those referred to _Mantellia_, he
named _Bennettites portlandicus_. There is no adequate reason for the
retention of the generic name _Mantellia_. The close resemblance of the
short and thick stems (12–30 cm. in height) described by Buckland to
those of some recent Cycads was recognised by Robert Brown and Loddiges
and the former suggested to Buckland the inclusion of the fossils in
a new family Cycadeoideae. It was this suggestion that led Buckland
to adopt _Cycadeoidea_ as a generic name. In a later account of the
Portland stems Buckland[1078], in deference to Brongniart’s opinion,
substituted Brongniart’s name _Cycadites_ for _Cycadeoidea_. It is
noteworthy that no reference is made in the original description to
the occurrence of lateral shoots among the persistent petiole-bases
that encase the Portland stems, but in a later account such shoots
are represented in one of the figured stems and are compared with the
buds occasionally produced on stems of _Cycas_[1079]. The subsequent
researches of Carruthers[1080] demonstrated the reproductive nature
of precisely similar lateral shoots in the stem on which he founded
the genus _Bennettites_. The generic name _Echinostipes_ given by
Pomel[1081], who had a passion for instituting new nomenclature, to
Buckland’s Portland stems has not been adopted: his genus _Crossozamia_
proposed for certain stems and fronds was revived by Carruthers[1082]
but has not been generally used. A further complication in the
nomenclature of Cycadean stems was introduced by Saporta’s
institution[1083] of _Bulbopodium_ and _Cylindropodium_: in the former
genus he included the small ovoid stem figured by Lindley and Hutton
as _Cycadeoidea pygmaea_[1084] and two French Jurassic species which
might reasonably be assigned to _Cycadeoidea_. To _Cylindropodium_ were
referred some large French stems from Jurassic strata: an examination
of the type-specimens in Paris convinced me that they are typical
forms of _Cycadeoidea_. As Ward[1085] has pointed out, both Saporta’s
genera may be merged in _Cycadeoidea_. Fliche and Zeiller[1086] also
include the small globular stems named by Saporta _Bulbopodium_ in
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