Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
This practice I followed in 1895[1091], but
my usage of the two generic names has not been consistent, the name
_Cycadeoidea_ being afterwards employed[1092] in a more comprehensive
sense. The investigation of the reproductive shoots of American Lower
Cretaceous and Upper Jurassic Cycadean stems[1093] has demonstrated the
practical identity of their ovulate strobili with those of the English
_Bennettites_. It is clear that the American stems as well as numerous
specimens from Italy and other European localities are generically
inseparable from _Bennettites_. The adoption of _Cycadeoidea_ by
Wieland, following Ward, has naturally strengthened the claim of
Buckland’s genus at least so far as wide usage is concerned, and with
some reluctance I propose to fall in with this terminology and for
the sake of convenience to give up the use of _Bennettites_. Among
Mesozoic Cycadean stems agreeing with those of recent genera in habit
and in the presence of an armour of persistent leaf-bases are several
which afford no evidence either of the occurrence of fertile shoots
or of lateral shoots with terminal flowers. To such stems the name
_Cycadeoidea_ has been applied, the designation _Bennettites_ being
restricted to stems with lateral fertile shoots: this practice of
treating the absence of reproductive shoots as a characteristic feature
of _Cycadeoidea_ is regarded by Wieland as illogical and artificial;
and if, as seems probable, he is correct in believing that the majority
of the Mesozoic Cycadean plants culminated their vegetative period by
prolific development of flowers, the absence of flowering branches
would be contingent on a certain stage of development and not a generic
character. Dr Stopes[1094] has recently published facts with regard
to the anatomical features of Cycadean stems which have a bearing
on the vexed question of nomenclature, but are more important from
a morphological point of view as they indicate a closer agreement
between certain types of Lower Cretaceous stems and those of _Cycas_
and other recent genera, in which successive rings of vascular tissue
are developed, than has previously been recognised. In her diagnosis of
_Cycadeoidea_ she includes the following statement: ‘In its internal
anatomy the trunk shows two or more (up to eight are recorded) zones
of secondary wood, the zones composed of distinct series of tracheids
each more or less regularly arranged in radial sequence.’ It is
claimed that the addition of this character constitutes for the first
time a clear distinction between _Cycadeoidea_ and _Bennettites_. Dr
Stopes states that the two stems on which Buckland founded the genus
_Cycadeoidea_ are lost and adds that Buckland’s type has certain
anatomical features which are not found in _Bennettites_. Buckland
in his description of _C. microphylla_ states that there are ‘two
laminated circles’ in the stem instead of the usual single cylinder,
a character suggestive of _Cycas_. In the original figure of this
Public-domain text, read in full here on John Shaqi.
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