Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
the older genus _Cycadeoidea_: one such type from the Portlandian of
Boulogne is described by these authors as _Cycadeoidea pumila_. Saporta
also proposed the name _Platylepis_[1087], the type-species being
_Cycadeoidea micromyela_ Mor., from the Lias of Calvados, which has
recently been investigated by Lignier[1088] who wisely adopts Morière’s
designation. Another unnecessary generic name is _Schizopodium_ given
by Morière to a stem, _S. Renaulti_[1089], regarded by Lignier as
indistinguishable specifically from _C. micromyela_.
The two names _Cycadeoidea_ and _Bennettites_ have been used by
authors for stems which are unquestionably generically identical
and as is often the case much confusion has been caused through the
failure of palaeobotanists to arrive at an agreement. Lester Ward, a
staunch advocate of the rule of priority, repeatedly pointed out that
Buckland’s name _Cycadeoidea_ should take precedence of _Bennettites_
on the ground that stems for which these genera were instituted are
clearly of the same type, and he added that the older genus, though
abandoned by its author, cannot be given up without violating the
inexorable rules of priority. On the other hand it has been urged that
the genus _Bennettites_ as defined by Carruthers is characterised
by the possession of (i) fertile shoots bearing strobili and (ii)
vegetative organs exhibiting certain anatomical characters. Buckland’s
species of _Cycadeoidea_, though bearing lateral shoots, are not well
enough preserved to afford definite information as to the morphological
features of the strobili, nor have we satisfactory data with regard
to the degree of resemblance between the vegetative features of
_Bennettites_ and the Portland stems. Carruthers laid stress on the
elliptical section of the stele in _Bennettites_ in contrast to the
cylindrical cylinder of Buckland’s _Cycadeoidea_ trunks. Subsequent
research has shown that this difference is not in itself a valid
criterion either of generic or specific rank but, as Wieland says, it
is an open question to what extent the stem structure of _Cycadeoidea_
and _Bennettites_ agrees. The probability is that were our knowledge
of the Portland stems less incomplete, they would be found to possess
no anatomical features inconsistent with this generic union. Granting
the correctness of this view, the adoption of the later generic term
would be a reasonable course to follow on the ground that it stands
for stems showing well preserved structure, while _Cycadeoidea_ was in
the first instance applied to stems showing only partially preserved
stumps of strobili, and the name was not retained by its author. Graf
Solms-Laubach[1090] and some other authors have used _Bennettites_ for
Cycadean stems possessing strobili of known structure such as those of
_Bennettites Gibsonianus_, reserving _Cycadeoidea_ for similar stems
but without strobili sufficiently well preserved to afford evidence of
morphological characters.
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