Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
of the seeds of _N. heterophylla_ as _Neurospermum Kidstoni_. The
generic term _Neuropterocarpus_ used by Grand’Eury in 1904[325],
though not defined by him, has priority and avoids the adoption of a
new designation for seeds attached to _Neuropteris_ fronds. In his
definition of _Neurospermum_ Arber makes no reference to the obliquity
of the apical snout that is clearly shown in fig. 422. There is no
evidence that _Neuropterocarpus_ possessed a cupular investment
comparable with that of _Lagenostoma_. Additional instances of the
association of seeds with _Neuropteris_ fronds are recorded by Renier
from the Belgian Coal Measures, also by Bertrand and Chodat from
France: Renier found seeds associated with _N. Schlehani_ Stur and with
the same species of frond Bertrand found impressions of oval ribbed
seeds. Prof. Chodat[326] has figured some fragments of _Neuropteris_
pinnules referred to _N. auriculata_ Brongn. from the Stephanian
of France which show small seed-like bodies apparently in organic
connexion with the lamina; but the specimens are too imperfect to
afford any satisfactory evidence as to the nature of the reproductive
organs.
Lotsy[327] has expressed the opinion that the bodies attached to
_Neuropteris_ pinnae described by Kidston and other authors as seeds
may possibly be vegetative buds, but if this were the case one would
expect to find some evidence of the bud-nature in some at least of the
specimens that have already been found.
It would seem that the microsporophylls of _Neuropteris_ were
constructed on different plans, some being of the type described by
Zeiller and other observers as _Potoniea_, while others bore sporangia
on pinnae without any accompanying laminae; but our knowledge of
the latter form represented by Kidston’s specimen of _Neuropteris
heterophylla_ (fig. 421, D) is very incomplete. On the other hand the
seeds appear to have been characterised by features suggesting a close
affinity to _Trigonocarpus_ and pointing to membership of the same
family.
_Alethopteris._
Although no specimens have been discovered showing actual connexion
between fronds and seeds, it is practically certain that _Alethopteris_
leaves, or at least some species of the genus, bore seeds of the
_Trigonocarpus_ type. The association of _Trigonocarpus Parkinsoni_
with _Alethopteris lonchitica_ is too frequent to be fortuitous and
there is further evidence afforded by certain anatomical resemblances.
In France other species of _Alethopteris_, _e.g._ _Alethopteris Serlii_
and _A. Grandini_, occur in association with _Pachytesta_[328], a large
seed similar to _Trigonocarpus_, and _Trigonocarpus_ is found in the
Pas-de-Calais coal-field with _Alethopteris Serlii_.
=TRIGONOCARPUS.= Brongniart.
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