Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
The specimen (fig. 430) is 9 cm. long and 4·2 cm. broad: at the
slightly contracted and broken base is a piece of immature axis (fig.
430, B, _a_) 12 mm. in diameter overtopped by a mass of closely packed
leaves encircling one another like the bulb-scales of an Onion (fig.
430 A, C)[382]. Most of the leaves included in the bud were attached
to the axis below the broken base. The curved, dichotomously branched,
veins are seen on some of the pieces of lamina on the surface of
the bud (fig. 430, A). The considerable breadth of the leaves is
demonstrated by the longitudinal and transverse sections. In fig. C
most of the laminae can be traced through the whole height of each
of the steep-sided arches: a few overlapping margins are seen in
fig. D. The veins are for the most part imperfectly preserved and
appear as clear spaces at regular intervals in the brown mesophyll.
The axis of the shoot consists of homogeneous parenchyma except near
the sloping sides where narrow dark bands (fig. 430, B, _a_) mark
the position of desmogen-strands of thin-walled elongated elements
representing an early stage in the development of vascular bundles
some of which have already produced spiral tracheids. Short secretory
cells accompany the immature conducting elements. The lamina slightly
exceeds 2 mm. in thickness in the broadest part: the mesophyll is
composed of large parenchymatous cells of elliptical or spherical
form often loosely attached owing to the well-developed system of
intercellular spaces. The lower epidermis, assuming that the outer
face of the rolled leaves is the morphologically lower surface, forms
a uniform layer of palisade cells characterised by their free conical
ends (fig. 431, A, B, _e_) which in some oblique sections appear as
sharply pointed papillae with almost filiform apices; but while the
cells were doubtless papillose like those of the epidermis of a velvety
petal, the pointed form is due in part to the greater distinctness of
the dark contents as compared with the lighter cell-walls. The upper
epidermis is much less distinct; it consists of smaller flattened
cells with occasional stomata. Renault[383] figures a specimen with
stomata in a better state of preservation. The vascular bundles are
rendered conspicuous by large secretory cells on the lower side, in
the larger veins in the form of an arc or irregular group (fig. 432,
A), but in the finer veins as single cells (fig. 431, A, B). These
sacs resemble the tannin cells accompanying the veins in a leaf of
_Ginkgo_ (_cf._ fig. 631, G). The xylem-elements are of two kinds,
(i) elongated spiral and scalariform conducting elements, forming a
vertical plate of a few rows in the larger veins (figs. 431, 432) or a
small compact group in the more slender veins (fig. 413, A, B); (ii)
much larger isodiametric cells with reticulate or spiral thickening
resembling the transfusion-tracheids of Conifers or, perhaps more
closely, similar elements in the leaves of _Lepidodendron_. These short
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