Fossil plants, Vol. 3 : $b A text-book for students of botany and geologySeward, A. C. (Albert Charles)
Science
Fossil plants, Vol. 3 : $b A text-book for students of botany and geology
Seward, A. C. (Albert Charles)
Paleobotany
tracheids are especially abundant on the flanks of the conducting
tracheids (figs. 431, _t_; 432, A, _t_), but they sometimes form a
complete investment. In the obliquely cut vein reproduced in fig. 431,
D, the transfusion-tracheids are abundant: a few are enlarged in fig.
431, E. In the smaller veins (fig. 431, A, B) they are represented by
the larger elements, _t_, on the sides of the conducting strands. The
protoxylem lies close to the upper edge in the middle line (_px_, figs.
431, A; 432, A); it is difficult to determine its precise position,
but it would seem to be slightly internal, the bundle being not quite
endarch. No phloem was recognised in the British Museum specimen, but
it presumably occurred, if present, where the black patch is shown
in fig. 432, A. Renault describes some phloem in sections which he
examined.
[Illustration: Fig. 431. _Dolerophyllum Goepperti_. Transverse
sections of leaves. _e_, epidermis; _s_, secretory cells; _t_,
transfusion-tracheids; _px_, protoxylem. (British Museum.)]
The mesophyll next the upper surface is in most cases represented
by spaces between the veins which give a crenulated outline to the
parenchyma (fig. 430, C, D); in some places the spaces contain remains
of very loose and crowded cells suggesting the original presence of
very lacunar tissue or possibly of thin-walled storage-cells. The
confinement of stomata to what is assumed to be the upper surface may,
as Renault and others have suggested, indicate leaves which floated on
water, an inference opposed to the view that the gaps in the mesophyll
mark the position of water-tissue.
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