The shape and structure of the zooecium differs greatly in different
groups of polyzoa. In its simplest form it is merely a cylindrical tube
of living matter which secretes an outer horny or gelatinous covering.
It is open at the end furthest from its base, at which it is attached
either to another zooecium or to some kind of supporting structure.
Certain parts of the polypide can always be extruded from the aperture,
which is known technically as the "orifice," or withdrawn through it
into the zooecium. When the polypide is retracted it draws in with it a
portion of the zooecium. The dead outer layer or ectocyst lines part of
the portion thus invaginated and forms the walls of a cavity within the
orifice. The base of this cavity consists in many forms of a transverse
partition pierced in the middle by a circular hole and known as the
"diaphragm." The diaphragm, however, does not constitute the limit of
the invaginated portion of the zooecium, for the living inner wall or
endocyst is dragged in still further and forms a sheath round the
retracted tentacles. When the tentacles are protruded they emerge
through the hole in the diaphragm, carrying with them their sheath of
endocyst. The invagination above the diaphragm, consisting of both
endocyst and ectocyst, is then everted.
The tentacles are a characteristic feature of the polypide. Together
with the base to which they are attached they are known as the
"lophophore"; they surround the mouth, usually in a circle. They differ
widely from the tentacles of _Hydra_ in both structure and function,
although they too serve as organs for the capture of prey; they are not
highly contractile and are not provided with nettle-cells but are
covered with cilia, which are in constant motion. When extruded they
form a conspicuous calix-like crown to the zooecium, but in the
retracted condition they are closely pressed together and lie parallel
to one another. They are capable individually of motion in all
directions but, although they usually move in concert, they cannot as a
rule seize objects between them.
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