The mouth is a hole situated in the midst of the tentacles. It leads
directly into a funnel-shaped oesophagus, the upper part of which is
lined with cilia and is sometimes distinguished as the "pharynx," while
the lower part, the oesophagus proper, is a thin-walled tube that
connects the pharynx with the stomach, which it enters on the dorsal
side. The stomach is a bulky organ that differs markedly in form and
structure in different groups of polyzoa. It is lined internally with
glandular cells and the inner wall is sometimes thrown into folds or
"rugæ." The part with which the oesophagus communicates is known as the
"cardiac" portion, while the part whence the intestine originates is
called the "pylorus" or "pyloric" portion. The intestine commences on
the ventral side opposite the entrance of the oesophagus and nearly on a
level with it, the bulk of the stomach depending between the two tubes.
This part of the stomach is often produced into a blind tube, the fundus
or cæcum. The alimentary canal may therefore be described as distinctly
Y-shaped. The proximal part of the intestine is in some forms lined with
cilia, and the tube as a whole is usually divided into two parts--the
intestine proper, which is nearest the stomach, and the rectum, which
opens by the anus not far from the mouth.
The nervous system consists of a central ganglion or brain, which is
situated at the base of the tentacles on the side nearest the anus and
gives out radiating nerves in all directions. Close to the brain and
providing a communication between the cavity of the zooecium and the
cavity in which the tentacles are contained (or, in the case of an
expanded polyp, the external world) is a ciliated tube known as the
"intertentacular organ." Apparently it acts as a passage through which
the genital products are expelled; but contradictory statements have
been made regarding it, and perhaps it is present only at certain
seasons or in certain conditions of the polypide.
[Illustration: Fig. 30.--Vertical section through a polypide of
_Alcyonidium_ with the polypide retracted (after Prouho).
A=orifice; B=contracted collar; C=diaphragm; D=parieto-vaginal muscles;
E=tentacles; F=pharynx; G=oesophagus; H=stomach; J=intestine; K=rectum;
L=intertentacular organ; M=retractor muscle; N=testes; O=ovary;
P=funiculus; Q=parietal muscles; R=ectocyst; S=endocyst.]
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