The muscular system is often of a complicated nature, but three sets of
muscles may be distinguished as being of peculiar importance, viz., (i)
the retractor muscles, which are fixed to the base of the lophophore at
one end and to the base of the zooecium at the other, and by contracting
pull the former back into the zooecium; (ii) the parieto-vaginal
muscles, which connect the upper part of the invaginated portion of the
zooecium with the main wall thereof; and (iii) the parietal muscles,
which run round the inner wall of the zooecium and compress the zooecium
as a whole. The parietal muscles are not developed in the
Phylactolæmata, the most highly specialized group of freshwater polyzoa.
The cavity between the polypide and the zooecium contains a reticulate
tissue of cells known as the "funicular" tissue, and this tissue is
usually concentrated to form a hollow strand or strands ("funiculi")
that connect the outer wall of the alimentary canal with the endocyst.
This rapid sketch of the general anatomy of a simple polyzoon will be
the best understood by comparing it with fig. 30, which represents, in a
somewhat diagrammatic fashion, a vertical section through a single
zooecium and polypide of the order Ctenostomata, to which some of the
freshwater species belong. The polypide is represented in a retracted
condition in which the Y-shaped disposition of the alimentary canal is
somewhat obscured.
In the great majority of cases the polyzoa form permanent colonies or
polyparia, each of which consists of a number of individual zooecia and
polypides connected together by threads of living tissue. These colonies
are formed by budding, not by independent individuals becoming
associated together. In a few cases compound colonies are formed owing
to the fact that separate simple colonies congregate and secrete a
common investment; but in these cases there is no organic connection
between the constituent colonies. It is only in the small subclass
Entoprocta, the polypides and zooecia of which are not nearly so
distinct from one another as they are in other polyzoa (the Ectoprocta),
that mature solitary individuals occur.
As representatives of both subclasses of polyzoa and of more than one
order of Ectoprocta occur in fresh water, I have prefaced my description
of the Indian species with a synopsis of the more conspicuous characters
of the different groups (pp. 183-186).
CAPTURE AND DIGESTION OF FOOD: ELIMINATION OF WASTE PRODUCTS.
The food of all polyzoa consists of minute living organisms, but its
exact nature has been little studied as regards individual species and
genera. In _Victorella bengalensis_ it consists largely of diatoms,
while the species of _Hislopia_ and _Arachnoidea_ possess an alimentary
canal modified for the purpose of retaining flagellate organisms until
they become encysted. Similar organisms form a large part of the food of
the phylactolæmata.
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