Most of the Gymnolæmata are marine, but species belonging to two of the
three suborders into which they are divided often stray into brackish
water, while a few genera that belong to one of these two suborders are
practically confined to fresh water. The three suborders are
distinguished as follows:--
Suborder A. _CHEILOSTOMATA._
The zooecia are provided with a "lip" or lid hinged to the posterior
margin of the orifice (see fig. 33, p. 175). This lid closes
automatically outside the zooecium or in a special chamber on the
external surface (the "peristome") when the polypide retracts and is
pushed open by the tentacles as they expand. The majority of the zooecia
in each zoarium are more or less distinctly flattened, but some of them
are often modified to form "vibracula" and "avicularia."
The Cheilostomata are essentially a marine group, but some species are
found in estuaries and even in pools and ditches of brackish water (fig.
33).
Suborder B. _CTENOSTOMATA._
The zooecia are provided with a collar-like membrane which is pleated
vertically and closes together above the polypide inside the zooecium
when the former is retracted; it is thrust out of the zooecium and
expands into a ring-shaped form just before the tentacles are extruded.
The zooecia are usually more or less tubular, but in some genera and
species are flattened.
The majority of the Ctenostomata are marine, but some genera are found
in estuaries, while those of one section of the suborder live almost
exclusively in fresh water.
Suborder C. _CYCLOSTOMATA._
The zooecia are provided neither with a lip nor with a collar-like
membrane. They are tubular and usually have circular orifices.
The Cyclostomata are exclusively marine.
Order II. PHYLACTOLÆMATA.
Ectoproctous polyzoa the polypides of which have a leaf-shaped organ
called an epistome projecting upwards and forwards within the circle of
tentacles and between the mouth and the anus. The zooecia are not
distinct from one another, but in dendritic forms the zoarium is divided
irregularly by chitinous partitions.
The Phylactolæmata are, without exception, freshwater species.
II.
SYNOPSIS OF THE LEADING CHARACTERS OF THE DIVISIONS OF THE SUBORDER
CTENOSTOMATA.
Suborder B. _CTENOSTOMATA._
The suborder has been subdivided in various ways by different authors.
The system here adopted is essentially the same as that proposed in a
recent paper by Waters (Journ. Linn. Soc. London, Zool. xxi, p. 231,
1910), but I have thought it necessary to add a fourth division to the
three adopted by that author, namely, the Alcyonellea, Stolonifera, and
Vesicularina. This new division includes all the freshwater genera and
may be known as the Paludicellina. In none of these divisions are the
tentacles webbed at the base.
The four divisions may be recognized from the following synopsis of
their characteristic features:--
Division I. ALCYONELLEA.
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