Most sponges possess a supporting framework or skeleton. In some it is
formed entirely of a horny substance called spongin (as in the
bath-sponge), in others it consists of spicules of inorganic matter
(either calcareous or siliceous) secreted by special cells, or of such
spicules bound together by spongin. Extraneous objects, such as
sand-grains, are frequently included in the skeleton. The spongin is
secreted like the spicules by special cells, but its chemical structure
is much more complicated than that of the spicules, and it is not
secreted (at any rate in most cases) in such a way as to form bodies of
a definite shape. In the so-called horny sponges it resembles the chitin
in which insects and other arthropods are clothed.
* * * * *
In no adult sponge do the collar-cells completely cover the whole of the
internal surface, the olynthus being a larval form, and by no means a
common larval form. It is only found in certain sponges with calcareous
spicules. As the structure of the sponge becomes more complicated the
collar-cells are tucked away into special pockets or chambers known as
ciliated chambers, and finally the approach to these chambers, both from
the external surface and from the inner or gastral cavity, takes the
form of narrow tubes or canals instead of mere pores. With further
complexity the simple internal cavity tends to disappear, and the sponge
proliferates in such a way that more than one osculum is formed. In the
class Demospongiæ, to which the sponges described in this volume belong,
the whole system is extremely complicated.
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