The colouring matter produced by the sponge itself is of two
kinds--pigment, which is probably a waste product, and the substances
produced directly by the ingestion of food or in the process of its
digestion. When pigment is produced it takes the form of minute granules
lying in the cells of the parenchyma, the dermal membrane being as a
rule colourless. Very little is known about the pigments of freshwater
sponges, and even less about the direct products of metabolism. It is
apparently the latter, however, that give many otherwise colourless
sponges a slight pinkish or yellowish tinge directly due to the presence
in cells of the parenchyma of minute liquid globules. In one form of
_Spongilla carteri_ these globules turn of a dark brown colour if
treated with alcohol. The brilliant colour of _Pectispongilla aurea_ is
due not to solid granules but to a liquid or semi-liquid substance
contained in the cells.
The green corpuscles of the Spongillidæ are not present in all species.
There is every reason to think that they represent a stage in the
life-history of an alga, and that they enter the sponge in an active
condition (see p. 49).
A fourth cause for the coloration of freshwater sponges may be noted
briefly. It is not a normal one, but occurs commonly in certain forms
(e. g., _Spongilla alba_ var. _bengalensis_). This cause is the growth
in the canals and substance of the sponge of parasitic algaæ, which turn
the whole organism of a dull green colour. They do not do so, however,
until they have reduced it to a dying state. The commonest parasite of
the kind is a filamentous species particularly common in brackish water
in the Ganges delta.
EXTERNAL FORM AND CONSISTENCY.
[Illustration: Fig. 5.--Part of a type-specimen of _Spongilla lacustris_
subsp. _reticulata_ (nat. size).]
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