I do not know exactly why light and shade must be mixed in a habitat
favourable for the growth of sponges, for most species prefer shade, if
it be not too dense; but it is certainly the case that, with a few
exceptions, Indian Spongillidæ flourish best in water shaded at the
edges by trees and exposed to sunlight elsewhere. One of the exceptions
to this rule is the Indian race of _Spongilla lucustris_, which is found
in small pools of water in sand-dunes without a particle of shade.
Several species are only found on the lower surface of stones and roots
in circumstances which do not suggest that their position merely
protects them from mud, which, as Mr. Potts points out, is their "great
enemy." A notable instance is _Trochospongilla pennsylvanica_, which is
found hiding away from light in America and Europe as well as in India.
It is curious that it should be easy to exterminate the sponges in a
pond by cleaning it out, for one would have thought that sufficient
gemmules would have remained at the edge, or would have been brought
rapidly from elsewhere, to restock the water. Mr. Green has, however,
noted that _Spongilla carteri_ has disappeared for some years from a
small lake at Peradeniya in which it was formerly abundant, owing to the
lake having been cleaned out, and I have made similar observations on
several occasions in Calcutta.
The question of the objects to which sponges attach themselves is one
intimately connected with that of the injury done them by mud. The delta
of the Ganges is one of the muddiest districts on earth. There are no
stones or rocks in the rivers and ponds, but mud everywhere. If a sponge
settles in the mud its canals are rapidly choked, its vital processes
cease, and it dies. In this part of India, therefore, most sponges are
found fixed either to floating objects such as logs of wood, to vertical
objects such as the stems of bulrushes and other aquatic plants, or to
the tips of branches that overhang the water and become submerged during
the "rains." In Calcutta man has unwittingly come to the assistance of
the sponges, not only by digging tanks but also by building
"bathing-ghats" of brick at the edge, and constructing, with æsthetic
intentions if not results, masses of artificial concrete rocks in or
surrounding the water. There are at least two sponges (the typical form
of _Spongilla alba_ and _Ephydatia meyeni_) which in Calcutta are only
found attached to such objects. The form of _S. alba_, however, that is
found in ponds of brackish water in the Gangetic delta has not derived
this artificial assistance from man, except in the few places where
brick bridges have been built, and attaches itself to the stem and roots
of a kind of grass that grows at the edge of brackish water. This sponge
seems to have become immune even to mud, the particles of which are
swallowed by its cells and finally got rid of without blocking up the
canals.
Public-domain text, read in full here on John Shaqi.
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