Fungi: Their Nature and UsesCooke, M. C. (Mordecai Cubitt)
Science
Fungi: Their Nature and Uses
Cooke, M. C. (Mordecai Cubitt)
Fungi
TREMELLINI.--In this order we have a great departure from the
character of the substance, external appearance, and internal
structure of the other orders in this family. Here we have a
gelatinous substance, and the form is lobed, folded, convolute, often
resembling the brain of some animal. The internal structure has been
specially illustrated by M. Tulasne,[N] through the common species,
_Tremella mesenterica_. This latter is of a fine golden yellow colour,
and rather large size. It is uniformly composed throughout of a
colourless mucilage, with no appreciable texture, in which are
distributed very fine, diversely branched and anastomosing filaments.
Towards the surface, the ultimate branches of this filamentous network
give birth, both at their summits and laterally, to globular cells,
which acquire a comparatively large size. These cells are filled with
a protoplasm, to which the plant owes its orange colour. When they
have attained their normal dimensions, they elongate at the summit
into two, three, or four distinct, thick, obtuse tubes, into which the
protoplasm gradually passes. The development of these tubes is unequal
and not simultaneous, so that one will often attain its full
dimensions, equal, perhaps, to three or four times the diameter of the
generative cell, whilst the others are only just appearing. By
degrees, as each tube attains its full size, it is attenuated into a
fine point, the extremity of which swells into a spheroidal cell,
which ultimately becomes a spore. Sometimes these tubes, or spicules,
send out one or two lateral branches, each terminated by a spore.
These spores (about ·006 to ·008 _mm._ diameter) are smooth, and
deposit themselves, like a fine white dust, on the surface of the
_Tremella_ and on its matrix. M. Léveillé[O] was of opinion that the
basidia of the Tremellini were monosporous, whilst M. Tulasne has
demonstrated that they are habitually tetrasporous, as in other of the
Hymenomycetes. Although agreeing in this, they differ in other
features, especially in the globose form of the basidia, mode of
production of the spicules, and, finally, the division of the basidia
into two, three, or four cells by septa which cut each other in their
axis. This division precedes the growth of the spicules. It is not
rare to see these cells, formed at the expense of an unilocular
basidium, become partly isolated from each other; in certain cases
they seem to have separated very early, they then become larger than
usual, and are grouped on the same filament so as to represent a kind
of buds. This phenomenon usually takes place below the level of the
fertile cells, at a certain depth in the mucous tissue of the
_Tremella_.
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