Fungi: Their Nature and UsesCooke, M. C. (Mordecai Cubitt)
Science
Fungi: Their Nature and Uses
Cooke, M. C. (Mordecai Cubitt)
Fungi
MUCEDINES.--Here, on the other hand, the threads, if coloured at all,
are still delicate, more flexuous, with much thinner walls, and never
invested with an external cortical layer. One of the most important
and highly developed genera is _Peronospora_, the members of which are
parasitic upon and destructive of living vegetables. It is to this
genus that the mould of the too famous potato disease belongs.
Professor De Bary has done more than any other mycologist in the
investigation and elucidation of this genus; and his monograph is a
masterpiece in its way.[o] He was, however, preceded by Mr. Berkeley,
and more especially by Dr. Montagne, by many years in elucidation of
the structure of the flocci and conidia in a number of species.[p] In
this genus, there is a delicate mycelium, which penetrates the
intercellular passages of living plants, giving rise to erect branched
threads, which bear at the tips of their ultimate ramuli, sub-globose,
ovate, or elliptic spores, or, as De Bary terms them--conidia. Deeply
seated on the mycelium, within the substance of the foster plant,
other reproductive bodies, called oogonia, originate. These are
spherical, more or less warted and brownish, the contents of which
become differentiated into vivacious zoospores, capable, when
expelled, of moving in water by the aid of vibratile cilia. A similar
structure has already been indicated in _Cystopus_, otherwise it is
rare in fungi, if the _Saprolegniei_ be excluded. In _Botrytis_ and in
_Polyactis_, the flocci and spores are similar, but the branches of
the threads are shorter and more compact, and the septa are more
common and numerous; the oogonia also are absent. De Bary has selected
_Polyactis cinerea_, as it occurs on dead vine leaves, to illustrate
his views of the dualism which he believes himself to have discovered
in this species. "It spreads its mycelium in the tissue which is
becoming brown," he writes, "and this shows at first essentially the
same construction and growth as that of the mycelium filaments of
_Aspergillus_." On the mycelium soon appear, besides those which are
spread over the tissue of the leaves, strong, thick, mostly
fasciculate branches, which stand close to one another, breaking forth
from the leaf and rising up perpendicularly, the conidia-bearers. They
grow about 1 _mm._ long, divide themselves, by successively rising
partitions, into some prominent cylindrical linked cells, and then
their growth is ended, and the upper cell produces near its point
three to six branches almost standing rectangularly. Of these the
under ones are the longest, and they again shoot forth from under
their ends one or more still shorter little branches. The nearer they
are to the top, the shorter are the branches, and less divided; the
upper ones are quite branchless, and their length scarcely exceeds the
breadth of the principal stem. Thus a system of branches appears, upon
which, on a small scale, a bunch of grapes is represented. All the
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