Geographic Variation in Red-backed Mice (Genus Clethrionomys) of the Southern Rocky Mountain Region — John Shaqi
Geographic Variation in Red-backed Mice (Genus Clethrionomys) of the Southern Rocky Mountain RegionCockrum, E. Lendell
Science
Geographic Variation in Red-backed Mice (Genus Clethrionomys) of the Southern Rocky Mountain Region
Cockrum, E. Lendell
Animals -- Variation; Mice
_Remarks._--Merriam (N. Amer. Fauna, 5:119, July 30, 1891) named this
subspecies on the basis of two specimens collected in the Black Hills
of South Dakota in July, 1888, and assigned it to the species
_Evotomys_ [= _Clethrionomys_] _gapperi_. He reported the diagnostic
characteristics as: "Similar to _E. gapperi_, but with larger ears and
shorter tail. The hazel of the dorsal area is not so bright as in
_gapperi_; the sides are the same golden brown." Of the cranial and
dental characteristics he wrote: "Much as in _E. gapperi_."
Bailey (Proc. Biol. Soc. Washington, 11:129, May 13, 1897), in his
"Revision of the American voles of the genus _Evotomys_," with one
additional specimen available, raised the Black Hills population to
specific status, re-emphasizing the shortness of the tail, and
pointing out a few slight cranial differences ("zygomatic arches low
and flaring out, so that the inner instead of the outer side shows in
top view; auditory bullae as large as in _gapperi_, but less
rounded").
Bailey (_loc. cit._) remarked that: "though based on so scanty
material, the characters distinguishing the species are fairly
pronounced. Its range is isolated and widely separated from that of
any other members of the genus by open prairie country and a wide belt
of the Transition zone. There seems to be no valid reason for
considering it a subspecies."
Additional specimens have been taken in recent years from the Black
Hills of South Dakota and northeastern Wyoming. This material has shed
light on the relationships and morphological characteristics of the
red-backed mice of this region. Bole and Moulthrop (Sci. Publ.
Cleveland Mus. Nat. Hist., 5:153, September 11, 1942) listed, as
comparative material, eight specimens from Bull Springs, Custer
County, South Dakota, under the name _Clethrionomys gapperi
brevicaudus_ (Merriam). They gave no reason for arranging
_brevicaudus_ as a subspecies of _C. gapperi_.
Twenty adults (11 skins and skulls, 9 skulls only) from Pennington
County, South Dakota (specimens in the University of Michigan
Museum of Zoology), have the following measurements (averages of
external measurements based on 11 specimens only): Total length, 142
(123-155); tail, 35 (30-39); hind foot, 19.5 (18.6-21.0); basal
length, 23.3 (21.7-24.5); condylobasilar length, 23.3 (21.9-24.5);
zygomatic breadth, 13.7 (12.9-14.7); lambdoidal breadth, 11.7
(11.3-12.9); alveolar length upper cheek-teeth, 5.5 (5.2-5.8);
interorbital breadth, 3.9 (3.6-4.1); length of nasals, 7.7 (7.1-8.5);
breadth of rostrum, 3.2 (2.9-3.6); and length of incisive foramina,
5.0 (4.6-5.3).
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