The first result was that the calcispongiæ afforded a splendid proof of
the impossibility of drawing sharp limits between species in the
perpetually developing animal world. In their case the different
varieties passed constantly out of each other and back into each other
in a way that would have made a classifier of the old type distracted.
But Haeckel had travelled far beyond the position of his boyhood, when
he had timorously concealed the bad species that would not fit into the
system. He said humorously that in the case of the calcispongiæ you had
the choice of distinguishing one genus with three species, or three
genera with 239 species, or 113 genera with 591 species. All this
confusion was saved by the Darwinian idea of not setting up absolutely
rigid classes, families, genera, and species. But even this was not yet
the essential point.
[Illustration:
ERNST HAECKEL, 1874.
]
As he had done in the case of the siphonophores, Haeckel endeavoured to
derive as much information as possible from the “ontogeny,” or embryonic
development, of the calcispongiæ. He established in some cases, it
seemed to him, that a single calcisponge-individual at first and up to a
certain stage developed from the ovum in the same way as a medusa or a
coral or an anemone. The fertilised ovum, a single cell, divided into
two cells, then several, and at last formed a whole cluster of cells. In
this cluster the cells arranged themselves at the surface, and left a
hollow cavity within. Then two layers of cells were formed, like a
double skin, in the wall of this vesicle, and an opening was left at one
spot in the wall of it. Thus we got a free-swimming embryo, with a
mouth, an external skin, and an internal digestive skin or membrane.
Then the creature attaches itself to the floor of the sea and becomes a
real sponge, partly by developing along its characteristic lines, and
partly (in most cases) by producing other sponges from itself in the
form of buds, like the siphonophore, and so forming an elaborate colony,
to which we give collectively the title of “a sponge.” These facts led
to the following reflections.
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