This original development from the ovum, first into an embryo with the
form of a small globe or, more correctly, an oval body consisting of two
layers of cells and having a hole at one pole—in other words, a creature
with nothing but skin, stomach, and mouth—was found, curiously enough,
in other animals besides the medusæ, corals, and sponges. We have the
same course of development in representatives of the most varied groups
of animals. There are worms, star-fishes, crabs, and snails that develop
in the same way. In fact, it was proved in this very year (1867) that
the lowest of the vertebrates, the amphioxus (or lancelet), develops in
the same way. And this was not all. In the ontogeny of all the higher
animals right up to man (inclusive) we find a state of things that most
closely resembles the same development. At all events, the fertilised
ovum gives rise in all cases to a cluster of cells; this cluster forms
something like a flattened or elongated vesicle with a single-layered
wall; the single layer of cells is doubled, and in the building up of
the body one half makes the external coat or skin and the other half the
internal lining or membrane. Haeckel reflected on the whole of the
facts, and drew his conclusions. This very curious agreement in the
earlier embryonic forms must be interpreted in terms of the biogenetic
law. In the case of the higher animals the forms have been profoundly
modified by cenogenesis. In the lower animals they are almost or
altogether a pure recapitulation of the real primitive course of the
development of the animal kingdom. In the earliest times animals were
evolved in something like the following way. First, the primitive
unicellular protozoa came together and formed crude social bodies,
clusters of cells that kept together, but had no special division of
labour. As all the members in the cluster pressed to the surface, in
order to obtain their food, they came to form, not a solid mass of
cells, but a hollow vesicle with a wall of cells. Then the first
division of labour set in. Certain cells, those that were situated at
the anterior pole, and so were better placed to receive the floating
food as the animal moved along, became the eating-cells of the group;
they provided nourishment for the others, as the nutritious sap
circulated through all the cells in the cluster, as we find in the case
of the siphonophores. As these feeding-cells multiplied rapidly at the
fore part of the animal, a depression was formed at that pole of the
body. In the end the ball or vesicle was doubled in upon itself, until
it came to have the form of a cup with a double-layered wall. Externally
were the cells in the skin that effected movement and feeling, and
afforded protection; inside, forming the internal wall, were the eating-
or stomach-cells. An opening remained at the top—the opening of the cup
or vase-like body. The food entered by it: it was virtually the “mouth.”
Public-domain text, read in full here on John Shaqi.
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