The occipital centers which mediate sensations of color are of course most closely associated with those other centers (probably the parietal) which receive sensations from the eye-muscles and which, therefore, mediate sensations which furnish space and position to the sensations of mere color. Now it is these occipital centers, mediators of light-sensations merely, which the experiments have shown most specially to be anæsthetic. The discharge of such centers means particularly the passage of excitations on to the parietal localization-centers. There are doubtless other outlets, but these are the chief group. The movements, for instance, which activity of these cells produces, are first of all eye-movements, which have to be _directly_ produced (according to our present psychophysical conceptions) by discharges from the centers of eye-muscle sensation. The principal direction of discharge, then, from the color-centers is toward the localization-centers. Now the experiment with falsely and correctly localized after-images proves that before the anæsthesia all localization is with reference to the point of departure, while afterwards it is with reference to the final fixation-point. The transition is abrupt. During the anæsthesia, then, the mechanism of localization is suffering a readjustment. It is proved that during this interval of readjustment in the centers of eye-muscle sensation the way is closed to oncoming discharges from the color-centers; but it is certain that any such discharge, during this complicated process of readjustment, would take the localization-centres by surprise, as it were, and might conceivably result in untoward eye-movements highly prejudicial to the safety of the individual as a whole. The much more probable event is the following: