Histology of medicinal plantsMansfield, William James
Science
Histology of medicinal plants
Mansfield, William James
Botany, Medical; Plant anatomy
The character of the outer wall of the epidermal cells differs
greatly in different plants. In most cases the wall is smooth; senna
is an example of such leaves. In certain other leaves the wall
is rough, the roughness being in the form of striations. In some
cases the striations occur in a regular manner; belladonna leaf is
typical of such leaves. In other instances the wall is striated in
an irregular manner as shown in chirata epidermis. Very often an
epidermis is rough, but the roughness is not due to striations.
In these cases the epidermis is unevenly thickened, the thin
places appearing as slight depressions, the thick places as slight
elevations. Boldus has a rough, but not a striated surface.
=Surface deposits= are not of common occurrence in medicinal plants;
waxy deposits occur on the stem of sumac, on a species of raspberry,
on the fruit of bayberry, etc. Resinous deposits occur on the leaves
and stems of grindelia species, and on yerba santa.
In certain leaves there are two or three layers of cells beneath the
epidermis that are similar in structure to the epidermal cells. These
are called hypodermal cells, and they function in the same way as the
epidermal cells.
Hypodermal cells are very likely to occur on the margin of the leaf.
Uva-ursi leaf has a structure typical of leaves with hypodermal
marginal cells. Uva-ursi, like other leaves with hypodermal cells has
a greater number of hypodermal cells at the leaf margin than at any
other part of the leaf surface.
The cutinized walls of epidermal cells are stained red with saffranin.
TESTA EPIDERMIS
=Testa epidermal cells= form the epidermal layers of such seeds as
lobelia, henbane, capsicum, paprika, larkspur, belladonna, scopola,
etc.
In surface view the end walls are thick and wavy in outline;
frequently the line of union--middle lamella--of two cells is
indicated by a dark or light line, while in others the wall between
two cells appears as a single wall. The walls are porous or
non-porous, and the color of the wall varies from yellow to brown, to
colorless. These cells always occur in masses, composed partially of
entire and partially of broken fragments.
In lobelia seed (Plate 4, Fig. 2) the line of union of adjacent cell
walls appears as a dark line. The walls are wavy in outline, of a
yellowish-red color and not porous.
In henbane seed (Plate 4, Fig. 3) the line of union between the cells
is scarcely visible; the walls are decidedly wavy, more so than in
lobelia, and no pits are visible.
In capsicum seed (Plate 4, Fig. 1) the cells are very wavy and
decidedly porous, the line of union between the cell walls being
marked with irregular spaces and lines.
In belladonna seed (Plate 5, Fig. 1) the walls between two adjacent
cells are non-striated and non-porous, and extremely irregular in
outline.
[Illustration: PLATE 4
TESTA EPIDERMAL CELLS
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