Histology of medicinal plantsMansfield, William James
Science
Histology of medicinal plants
Mansfield, William James
Botany, Medical; Plant anatomy
Collenchyma cells differ from parenchyma cells in a number of
ways: first, the cell cavity is smaller; secondly, the walls are
thicker, the greater amount of thickening being at the angles of the
cells--that is, the part of the cell wall which is opposite the usual
intercellular space of parenchyma cells, while the wall common to
two adjoining cells usually remains unthickened. In horehound stem
(Plate 30, Fig. 2) the thickening is so great at the angles that no
intercellular space remains. In the side column of motherwort stem
(Plate 30, Fig. 1) the thickening between the cells has taken place
to such an extent that the cell cavities become greatly separated and
arranged in parallel concentric rows.
The collenchyma of the outer angle of motherwort stem (Plate 30, Fig.
3) is greatly thickened at the angles. There are no intercellular
spaces between the cells, and cell cavity is usually angled in
outline instead of circular, as in the cells of horehound. In certain
plants intercellular spaces occur between the cells, and the walls
are striated instead of being non-striated, as in the stems of
horehound, motherwort, and catnip.
[Illustration: PLATE 29
_A._ Diagrammatic sketch of the cross-section of catnip stem
(_Nepeta cataria_, L.). 1. Collenchyma occurring at the four angles
of the stem.
_B._ Diagrammatic sketch of the cross-section of motherwort stem
(_Leonurus cardiaca_, L.). 1, 2, 3. Twelve masses of collenchyma
tissue occurring at the four sides of the stem.]
[Illustration: PLATE 30
COLLENCHYMA CELLS
1. Cross-section of a side column of the collenchyma of motherwort
stem (_Leonurus cardiaca_, L.).
2. Cross-section of the collenchyma of horehound stem (_Marrubium
vulgare_, L.).
3. Cross-section of the collenchyma of the outer angle of
motherwort stem.]
Collenchyma cells retain their living contents at maturity. Many
collenchyma cells, particularly of the outer layers of bark and the
collenchyma of the stems of herbs, contain chlorophyll.
The walls of collenchyma consist of cellulose.
STONE CELLS
=Stone cells=, like bast fibres, are branched or non-branched. Each
group is then separated into subgroups according to wall structure
(whether striated, or pitted and striated, etc.), thickness of wall
and of cell cavity, color of wall and of cell contents, absence of
color and of cell contents, etc.
BRANCHED STONE CELLS
=Branched stone cells= occur in a number of drugs. In witch-hazel
bark (Plate 31, Fig. 2) the walls are thick, white, and very porous.
In some cells the branches are of equal length; in others they are
unequal. In the tea-leaf (Plate 31, Fig. 1) the walls are yellowish
white and finely porous. When the lower wall is brought in focus, it
shows numerous circular pits. These pits represent the pores viewed
from the end. The branches frequently branch or fork.
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