Enough has been said to show that an important portion of the history
of phytotomy lies between the beginning and the end of the period under
consideration. The distance between the knowledge of the structure of
vegetable tissue which existed at the beginning of the century, and
that of Meyen and von Mohl on the same subject in 1840, is wonderfully
great; in the one case an uncertain groping about among obscure
ideas, in the other a complete exposition of the inner architecture
of the mature plant. But in spite of this great difference between
beginning and end, it is better to review the efforts of this period
of forty years as a connected process of historical development, and,
notwithstanding the interval between the appearance of Moldenhawer’s
contributions in 1812 and Meyen’s and von Mohl’s labours about 1840, to
consider the latter as the settlement of the questions taken up at the
commencement of the century. Moreover after 1840, with the appearance
of Schleiden and Nägeli on the scene, new points of view were suddenly
disclosed, and new aims were proposed in phytotomic investigation; it
is no objection to this view of the subject, that the most productive
portion of von Mohl’s labours falls in the succeeding twenty years, and
that during this later period his position is one of equal authority
with the new tendency and of participation in it. Up to 1845 his
discoveries were the culminating point of the older phytotomy; they put
the finishing stroke to the work which Mirbel, Link, Treviranus, and
Moldenhawer had begun. The object almost exclusively pursued during
all this period was to frame as true a scheme as possible of the inner
structure of the mature organs of the plant; it was requisite to
gain a right understanding of the diversities of cells and forms of
tissues, to classify them and supply them with names, and to secure
well-conceived definitions of these names. Hence almost exclusive
attention was paid to the configuration of the solid framework of
cell-membrane, and of this chiefly in the matured state, to the form
of the several elementary organs and their combination in the tissue,
to the sculpture of the wall-surfaces, and to the connection of
cell-spaces by pores or their separation by closed walls. There was
much discussion, especially at first, on the contents of vessels and
cells, and on supposed movements of sap in connection with anatomical
research, but there was no careful connected investigation of the
cell-contents; it was not yet recognised that the true living body of
the vegetable cell is only a definite part of the contents inclosed by
the cell-wall; the solid walls, the framework of the whole building,
were regarded as of primary importance in the structure of the cell.
It was not till the following period that in the light of historical
development another view asserted itself, namely, that the solid
framework of vegetable tissue with all its importance is yet in the
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