Bernhardi distinguishes vessels into two main groups, air-vessels and
vessels properly so called. He calls the first group air-vessels for
the same reason that led Grew to give them that name, namely, that they
are filled with air during a part at least of the period of vegetation;
they are found in the wood, and, where there is no closed woody body,
there the woody bundles are formed both of vessels and also of bast
strands which enclose vascular canals. These latter he next divides
into three chief kinds; annular vessels, which he was the first to
discover, true spiral vessels with a band which can be unwound, and
scalariform vessels, by which term he understood not only those with
broad slits, such as are found in Ferns, but also the pitted vessels in
secondary wood. His idea of annular and spiral vessels was perfectly
correct, and he mentions Hedwig’s notion already described, and shows
that its exact opposite is true, namely, that the spiral band is
surrounded by a membrane on the outside,—a fact which was afterwards
denied by Link, Sprengel, and Moldenhawer. On the other hand he did
not understand the sculpturing on the scalariform vessels; he took the
pits in the dotted vessels for thickenings of the wall, such as are
seen in the transverse ridges between the slits in true scalariform
vessels, and the slits he thought were closed. If there was much that
was erroneous in these views, yet Bernhardi contributed essentially
to the clearing up of the subject by his effort to distinguish the
different forms of air-vessels, and especially by pointing attention to
the fact that neither spiral nor annular vessels are found in secondary
wood. The resemblance between different forms of vessels misled many of
his contemporaries into supposing that they are due to metamorphosis
of true spiral vessels. Bernhardi showed that different forms of wall
are found inside one vascular tube, but that this does not depend on
modification with age; observation rather teaches that every kind of
vessel receives its character in its young state, and especially that
the youngest scalariform vessels do not present the form of spiral
vessels.
Under the head of vessels proper he reckoned all tubular forms filled
with a peculiar juice, milk-cells and true milk-vessels, and also
resin-ducts and the like, and he made many good and still valuable
observations on their distribution and sap-contents. He could not see
the differences of structure in these various fluid-conveying vessels
with the low magnifying power of his glass, and therefore attended
chiefly to the structure of the large resin-ducts, which on the whole
he rightly understood.
Public-domain text, read in full here on John Shaqi.
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