plants presents difficulties at the present day to a practised observer
with a good microscope. Occasionally observers had taken the stem of
the gourd, where the large cells and vessels suited small magnifying
power, but where many abnormal conditions occurred to interfere with
their conclusions. The Monocotyledons, like the Vascular Cryptogams,
had hitherto been comparatively neglected. When then Moldenhawer made
a monocotyledonous and rapidly growing plant, with very large-celled
tissue and comparatively very simple structure, the chief subject of
his investigations, he was sure to succeed in making out many things
more clearly than his predecessors. It was an important point that
he found the fibrous elementary organs in this plant united with
the vessels into bundles, which are separated by a strict line of
demarcation from the large-celled parenchyma that surrounds them. Thus
the peculiar character, the idea, of the vascular bundle was brought
prominently into contrast with that of other forms of tissue. This took
the place of the distinction between rind, wood, and pith, which had
served former phytotomists as the basis of their histological survey,
but which is in itself only a secondary result of the later elaboration
of certain parts of the plant. Moldenhawer, in laying the chief stress
from the first on the contrast between vascular bundles and parenchyma,
hit upon a histological fact of more fundamental importance, the right
appreciation of which has since enabled the phytotomist to find his way
through the histology of the higher plants. For while the construction
of Monocotyledons and Ferns must seem abnormal and quite peculiar to
any one who starts with examining the rind, wood, and pith of old
dicotyledonous stems, those on the contrary who, with Moldenhawer, have
recognised a special histological system in the vascular bundles of
Monocotyledons, have the way opened to them to seek for a similar one
in the Dicotyledons, and to refer the secondary phenomenon of wood and
rind to the primary existence of vascular bundles. Moldenhawer did in
fact open this way, when he showed how the growth of a dicotyledonous
stem may be understood from the structure and position of the
originally isolated vascular bundles (Beiträge, p. 49, etc.). But he
was thus of necessity led to the rejection of Malpighi’s theory of the
growth in thickness of woody stems, which all vegetable anatomists from
Grew to Mirbel had adopted. Though Bernhardi and Treviranus made weak
attempts to discredit it, Moldenhawer was the first who distinctly
rejected the origin of the external layers of wood from the inner bast,
and proposed the first really practical basis for the later and correct
theory of secondary growth in thickness (p. 35). The removal of this
ancient error is in itself a very important result, and one which,
apart from all other services, must secure him an honourable place in
the history of botany.
Public-domain text, read in full here on John Shaqi.
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