1. The view that the cell is the sole and fundamental element in
vegetable structure had been already maintained by Sprengel and Mirbel,
but not supported by exact observations. Treviranus too had shown
that the vessels in wood are formed by the union of rows of cell-like
tubes, but he had never arrived at a thoroughly clear conception of the
matter. On the one side was the theory that the plant consists entirely
of cells, on the other, and for long the old and strange view, that
the spiral thread was an independent elementary organ of vegetable
structure,—a view which Meyen still maintained in 1830. Von Mohl must
be regarded as the first who took up the all-important position, that
not only the fibrous elements of bast and wood, which had long been
considered to be elongated cells, but the vessels of the wood also
are formed from cells; and we may on this point give great weight to
his own assertion that he was the first who observed the formation
of vessels from rows of closed cells. This discovery happened in the
year 1831, and he describes distinctly, though briefly, the decisive
observations in his treatise on the structure of the palm-stem. At the
points of constriction in the vessels he saw the dividing walls, the
existence of which had been denied by all former phytotomists; ‘these
dividing walls,’ he says, ‘are entirely different from the rest of
the membranes of the plant, being formed of a network of thick fibres
with openings between them.’ He studied the history of the development
of these vessels both in palms and in dicotyledonous plants. ‘In the
young shoot,’ he says, ‘are found at the spots, where afterwards
there are large vessels, perfectly closed large cylindrical tubes
with a transparent and very delicate membrane.’ He then shows how by
degrees the sculpture peculiar to the walls of vessels is formed on the
inside of these tubes, and he takes the opportunity of saying that a
metamorphosis in time from one form of vessel into another is entirely
out of the question, as Treviranus also and Bernhardi had maintained.
‘The dividing walls (transverse septa),’ he continues, ‘are formed
in a precisely similar manner to the side-walls of vessels; only the
original tender membrane of the septa is usually lost in the meshes
of the network of fibres.’ From that time no phytotomist capable of
an independent judgment has had any doubt with regard to this view of
the formation of the vessels in wood. It is however striking enough
that von Mohl, who thought it so important to show that the cell is
the sole foundation of vegetable structure, never extended the proof
to milk-vessels and other secretion-canals in order to show whether
and how these also are formed from the cells. In his treatise on the
vegetable cell (1851) he still expressed doubt about Unger’s assertion,
that the milk-vessels are also formed from rows of cells that coalesce
with one another, and held rather to the view of an anonymous writer
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