in the ‘Botanische Zeitung’ of 1846, page 833, that these vessels are
membranous linings of gaps in the cell-tissue. He might well lose his
taste for the examination of these and similar organs after Schultz
Schultzenstein had by his various treatises, written after 1824, on the
so-called vital sap and the circulation which he attributed to it, made
this part of phytotomy a very quagmire of error, and had not refrained
from replying in an unbecoming manner to von Mohl, who repeatedly
opposed his views; moreover Schultz’s essay ‘Ueber die Circulation des
Lebenssafter’ (1833), which teems with absurdities, had received a
prize from the Academy of Paris.
2. The growth in thickness of the cell-membrane, and the sculpture
caused by it was a subject that is more or less connected with most
of von Mohl’s writings. He developed the chief features of his
view in 1828 in his first work, ‘Die Poren des Pflanzengewebes!’
The way in which he represented to himself the growth in thickness
of cell-membranes at a later time may be expressed as follows. All
elementary organs of a plant are originally very thin-walled perfectly
closed cells, which in the tissue are separated by walls formed of two
laminae[82]; on the inside of these primary cell-membranes, after they
have ceased to increase in circumference, new layers of membranous
substance are formed, which lying one upon another adhere closely
together, and represent the whole amount of secondary thickening
layers; on the inner side of the membrane thus thickened by apposition
there may usually[83] be perceived a tertiary layer of a different
character.
But there are certain sharply defined spots on the original cell-wall,
where this thickening does not take place; in such spots the cell is
still bounded only by the primary membrane; it is these thin spots
which bear the name of pits, and which Mirbel, and in some cases
Moldenhawer, took for holes, but von Mohl considered that it was only
in very exceptional cases that they were really changed into holes by
resorption of the thin primary wall. In accordance with this theory,
the spiral, annular, and reticulated vessels are produced by deposition
of thickening matter in the form suitable to each case on the inside
of the originally smooth thin cell-wall. But like Schleiden and other
phytotomists, von Mohl was not quite clear in his views either of the
origin or mode of formation of matured bordered pits; it was supposed
that the two laminae of the dividing wall parted from one another
at certain spots in such a manner that a lenticular hollow space
was formed between them, and that this space answered to the outer
border of the pit, while the inner border was the result of ordinary
pit-formation. This view, which could be shown to be incorrect by the
history of development, arose in fact from inexact observation,—a rare
case with von Mohl; the true course of events in the formation of
bordered pits was first described by Schacht in 1860.
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