5. Forms of tissue and comparative anatomy. Phytotomy up to 1830
had been weak in its classification of tissues, in its ideas as to
their arrangement, and consequently in its histological terminology;
the inconvenience arising from this state of things was most
distinctly felt when it became necessary to compare the structure of
different classes of plants, Cryptogams, Conifers, Monocotyledons
and Dicotyledons, and to establish their true differences and actual
agreements. How little phytotomy had advanced in this respect is
shown plainly in the account of tissues given by Meyen in his ‘Neues
System’ in 1837. To von Mohl belongs the merit of having perceived at
an early period in his scientific career, and more clearly than his
contemporaries, the value of a natural and sufficient discrimination
of the various forms of tissue, and the necessity of obtaining a
correct view of their relative disposition; he thus showed the way to
an understanding of the general structure of the higher plants, and
rendered it possible to make a scientific comparison of the structure
of different classes of plants.
Von Mohl, like Moldenhawer long before, showed from the first a correct
apprehension of the peculiar character of the vascular bundles as
compared with other masses of tissue. He, too, examined them first in
Monocotyledons, and gave an account of them in his treatise on the
structure of Palms (1831), and also in his later essays on the stems
of Tree-ferns, Cycads, and Conifers and on the peculiar form of stem
in Isoëtes and Tamus elephantipes, to be found in his ‘Vermischte
Schriften’ of 1845. His just conception of them as special systems
composed of various forms of tissue has made his account clear and
intelligible, and his whole treatment of the subject appears new in
comparison with that of every previous writer except Moldenhawer. If
these labours of von Mohl are surpassed in value by later studies of
the history of development, they served for the time as a nucleus
for further investigations, especially into the nature of stems. It
contributed in a high degree to a correct insight into the structure
of the stem, that von Mohl, agreeing in this with Moldenhawer,
distinguished the portion belonging to the wood from the portion
belonging to the bast in the vascular bundles, and regarded both as
essential constituents of a true vascular bundle. Not less important
were his enquiries into the longitudinal course of the vascular bundle
in the stem and leaf, which showed that in the Phanerogams the bundles
in the stem are only the lower extremities of the bundles, the upper
extremities of which bend outwards into the leaves, and that the
Monocotyledons and Dicotyledons agree in this particular, though the
course of the bundle differs considerably in the two cases. He obtained
an important result in this respect in his researches on palm-stems
in 1831, when he proved the incorrectness of the distinction between
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