cell-germs in the spores of Marchantia usually only from two to four
serve to form cells, the rest becoming overlaid with chlorophyll,
and being consequently withdrawn from the vital process. He who is
acquainted with the modern view of the processes of free cell-formation
founded on the numerous and careful investigations of later times will
scarcely discover in the above account of Schleiden’s theory a single
correct observation.
Soon after, von Mohl published in ‘Linnaea,’ 1839, p. 272, his
observations on the division of the mother-cells of the spores of
Anthoceros; these were carefully made and were correct in all the main
points; and in opposition to Mirbel’s former statements they establish
the fact, that the division is effected by the mucilaginous contents
of the cell, and consequently that it is not a passive division of
the contents of the mother-cell produced by the growth inwards of
projections of the cell-wall.
Unger[87] was the first to declare distinctly against Schleiden’s
doctrine, and his observations on the punctum vegetationis appeared
in the ‘Linnaea’ of 1841, p. 389; from the size and position of the
cells he concluded that the tissue-cells in this case are formed by
division, and not in the manner alleged by Schleiden. Soon after Nägeli
also (‘Linnaea,’ 1842, p. 252) observed the processes of cell-formation
in the extremities of roots, but he did not conceive them to be cases
of division; he saw two nuclei form in each mother-cell, and a new
cell form round each nucleus, and explained the origin of the dividing
wall as due to the meeting together of the two new cells; he thought
that a similar process takes place in stomata and in the mother-cells
of pollen; this conception was not absolutely incompatible with
Schleiden’s theory, but there was this difference, that in Nägeli’s
case essential processes were correctly observed, but were to some
extent incorrectly interpreted. In the same year appeared the first
edition of Schleiden’s ‘Grundzüge der wissenschaftlichen Botanik,’
in which his theory of cell-formation was repeated in a more precise
form. That he was thoroughly in earnest to maintain it is shown by the
fact that he gave still another exposition of it in his ‘Beiträge zur
Botanik’ in 1844, where he insists that his method of cell-formation
is the general one, though it has been distinctly ascertained in the
Phanerogams only. But how completely an observer may be led captive
by a preconceived opinion may be learnt from Schleiden’s suggestion,
that the formation of zygospores in Spirogyra is in accordance with his
views, though it is impossible to conceive of a case of cell-formation
more easy to observe, or less reconcilable with Schleiden’s theory.
It was mentioned in the first book, that Hedwig and Vaucher were
acquainted with the remarkable process of the formation of zygospores
in the alga-genus Spirogyra; but this as late as Schleiden’s time
was not regarded as an example of cell-formation, and his view was
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