We make acquaintance with Schleiden’s theory of cell-formation in its
original form, if we turn to his treatise, ‘Beiträge zur Phytogenesis,’
(in the Archiv für Anatomie, Physiologie, etc. von Johannes Müller,
1838). The work begins with some remarks on the general and fundamental
laws of human reason, etc., discusses the literature of cell-formation
in a few lines without mentioning von Mohl’s numerous observations,
goes on to mention the general occurrence of the nucleus which here
receives its new name, then occupies itself with gum, sugar, and
starch, and at last comes to the main subject. There are two spots,
says Schleiden, in the plant, where the formation of new organisation
may be most easily and most certainly observed, the embryo-sac and
the end of the pollen-tube, in the latter of which, according to his
theory of fertilisation, the first cells of the embryo are supposed to
be formed, but where in fact no cells are formed. At both spots small
granules soon arise in the gum-mucilage, which, before homogeneous,
now becomes turbid, and then single larger and more sharply defined
granules, the nucleoli, appear. Soon after, the cytoblasts are seen as
granular coagulations from the granular mass; they grow considerably in
this free condition, but as soon as they have reached their full size,
a delicate transparent vesicle is formed upon them; this is the young
cell, which at first presents the appearance of a very flat segment of
a sphere, whose plane side is formed by the cytoblast, the convex by
the young cell (the cell-membrane), which rests upon the cytoblast as
a watch-glass on a watch. Gradually the vesicle becomes larger and of
firmer consistence, and now the whole of the wall, except where the
cytoblast forms part of it, consists of a jelly. By-and-bye the cell
grows beyond the edge of the cytoblast and rapidly becomes so large
that the latter appears only as a small body inclosed in one of the
side walls. The shape of the cell becomes more regular with advancing
growth and under the pressure of adjoining cells, and often passes
into that of a rhombododecahedron, which Kieser for reasons drawn from
the nature-philosophy assumed to be the fundamental form. It is only
after the resorption of the cytoblast that the formation of secondary
deposits on the inner surface of the cell-wall commences, though some
exceptional cases are adduced. Schleiden thinks (p. 148) that he may
assume that the process here described is the general law of formation
of vegetative cell-tissue in Phanerogams. He adds particularly that the
cytoblast can never lie free inside the cell, but is always enclosed in
a duplication of the cell-wall, and he thinks that it is an absolute
law that every cell, except perhaps in cambium, begins as a minute
vesicle, and grows to the size which it reaches in its matured state.
The resemblance of this view to that of Sprengel and Treviranus is
increased by what we find further on, where we read that from the
Public-domain text, read in full here on John Shaqi.
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