We have already observed the influence of the theory of evolution on
the doctrine of the fertilisation of plants in the case of Morland
and Geoffroy. We learn more about it in the work, already quoted,
of the philosopher Christian Wolff, ‘Vernünftige Gedanken von den
Wirkungen der Natur,’ Magdeburg, 1723; it will be well to give his
own words, for they will serve to show at the same time the amount of
knowledge possessed by a cultivated and well-read man in the country
of Camerarius and thirty years after the appearance of his treatise
on the sexuality of plants. In the second chapter of the fourth part,
which treats of the life, death, and generation of plants, Wolff
says: ‘Ordinarily plants are produced from seeds, for the seed not
only contains the plant in embryo but also its first food.’ He says
that propagation by means of buds is equally natural, for each bud
contains a branch in little. ‘We find inside in the flower a number of
stalks disposed in a circle, and something at the top of each, which
is full of dust and lets the dust fall on the upper part of that which
holds the seed; this organ is compared by some to the genitals of the
animal, and the dust to the male seed; they think also that the seed
is made fruitful by the dust, and that therefore the embryo must be
conveyed by the dust into the seed-case and there be formed into seeds.
I have proposed to examine into the matter, but I have always let it
escape me.’ ... ‘Since all that has been hitherto adduced is found
also in flowers which spring from bulbs, and it is also certain that
the leaves of bulbs have consequently embryos in them ... it is easy
to see that the embryos must come from the leaves of the bulbs. And
since they could as easily be conveyed from there into the seed-grains
with the sap, as into the dust which is produced in the upper part of
the flower, I am inclined to think that this is the true account of
the matter and that it will be confirmed by experience. But now comes
the main question, whence come the embryos into the sap; since they
have not an external figure only but an internal structure also, it is
not plain how they can be formed either by the mere inner movement of
the sap, or by separation of certain parts.... And this is certainly
more credible, that the embryos already exist in little in the sap and
the plant, before they are brought by some change into the condition
in which they are met with in the seed and in buds. But there is the
further question where they were previously. They must either lie one
in another in a minute form, as Malebranche especially maintains, or
they are brought from the air and the earth with the nourishing sap
into the plant, an idea which Honoratus Fabri advanced and Perrault and
Sturm developed after him. According to the first opinion the first
seed-grain must have contained everything in itself, which has grown
from it to this hour.’ But this demand goes beyond even Wolff’s powers
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