of belief; for, says he, it is too great a tax on the imagination to
conceive of this inclosing of germs one in another like box within box.
It is well known that such notions as these were very prevalent in
the 18th century, and that the spermatozoids of animals were thought
to lend considerable support to them; even Albert Haller after 1760
was an adherent of the theory of evolution. However confused Wolff’s
general train of thought may be, we should notice his perception of
the fact, that the theory of evolution does away with the sexual
significance of the anthers. We shall see by-and-bye, that Koelreuter
was able to form a very different idea of sexual propagation. His great
importance in the history of the sexual theory will be best learnt
from a consideration of the speculative views of his predecessors
and contemporaries. It will not be amiss therefore to disregard
chronology for a while, and to notice here the views of the Baron von
Gleichen-Russworm, and the feeble arguments of Kaspar Friedrich Wolff
against the theory of evolution. The first-named writer in his work
‘Das Neueste aus dem Reich der Pflanzen,’ 1764, relying principally on
microscopic observation of the contents of pollen-grains, supported the
view that the granules in them answer to spermatozoids in animals, and
that they find their way into the ovule and are there developed into
embryos. Yet Gleichen was at the same time a zealous supporter of the
sexual theory, and endeavoured to meet well-known objections to it by
pointing to the occurrence of female flowers on male plants of spinach;
he also made some experiments on maize and hemp in the interests of
the theory. He did not perceive that hybrids supply convincing proof
against the theory of evolution, but he rightly appealed to them as
affording strong arguments in favour of sexuality. His real knowledge
of hybrids is partly drawn from the statements of Linnaeus, with
which we have already made acquaintance; he even describes a hybrid
between a goat and a cow, and other similar ones, and he is angry
with Koelreuter for fixing such narrow limits to the occurrence of
hybrids; thus the first person who produced hybrids systematically in
the vegetable kingdom must submit to be scolded for refusing to accept
the imaginary hybrids of his contemporaries. Gleichen’s book and the
selection from his microscopic discoveries, which appeared in 1777,
abound in good detached observations; he was the first who saw and
figured the pollen-tubes of Asclepias, without of course suspecting
their real nature and importance.
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