In the case of the Male Fern the germination of the spore will start in
about eight days, but in other species the period varies. Many of the
succeeding stages cannot be intelligently followed except with the aid of
a microscope. The first thing which happens to the germinating spore is
the development of a root hair which helps in fixing the process to the
soil. A system of cell division now commences in the other portion of the
spore which results in the formation of a green filament, every cell of
which is capable of producing root hairs. This filament is the beginning
of the body, known as the prothallus, which is responsible for the sex
organs. The process of cell-division goes forward and ultimately results
in the development of a green scale measuring, perhaps, an eighth of an
inch across at its broadest part. This is the fully-grown prothallus.
From the underside arise more root hairs, and it is here also that the
antheridia (male organs) and the archegonia (female organs) are produced.
As a rule, both kinds of organs occur on the same prothallus, although
now and again prothalli have been discovered which are exclusively male
or female. The matter is of interest, because it evidences an occasional
distinction of sex which has become habitual in some of the Club Mosses.
In the case of a perfectly normal prothallus the male organs or
antheridia are to be found amongst the root hairs, whilst the female
organs or archegonia arise from a cushion, several cells deep, more
towards the centre of the process.
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