The point may be best understood by considering the foot of man, of
which Fig. 60 shows a good example. The value of the roughened surface
of the foot with its papillary ridges can hardly have been great, even
in the days when man’s foot was naked, at any rate so little that for
him to acquire by a selectional process such a remarkable _change_ of
arrangement as we see when we look at the foot of man and of any other
Primate involves on our part a tremendous stretch of imagination as to
its _modus operandi_. These low, soft ridges of man’s foot could do
little to prevent him from slipping on such surfaces as grass, sand,
rock, wet or dry, and from the time when he began to protect his feet
with coverings this small value would be further reduced. _Underneath
his developing plantar arch it would not exist at all, and yet here
especially he has changed their direction._ As to the papillary
ridges, man’s foot has sadly embarked on the pathway of degeneration
much as his little toe has done. Not only has he here a much simpler
arrangement than any ape or monkey, but the individual ridges are
blurred and flattened on much of the plantar surface. This comes of
his pride in acquiring his human distinction, or title of nobility, of
a plantar arch and his coincident increase of pedestrian locomotion.
On the triple bases of support, heel, ball of great and little toe,
the ridges are still strongly marked and coarse; transverse on the
heel, whorl-like on the ball of the great toe, and oblique or nearly
transverse on that of the little toe. On the rest of the surface they
are vulgarly transverse. And I may add that the toe-prints of man are
simplicity itself compared with his finger-prints. It would seem that
this example of arrangement of ridges on man’s foot is strongly in
favour of the hypothesis that they are so disposed by flexion of the
foot in walking, and not by some need for prevention of slipping under
the guidance of selection.
Lower Animals.
[Illustration: Fig. 61.--Slow loris--right foot.]
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