At the other end of the scale the scanty ridges of a hedgehog’s or
squirrel’s foot would be negligible in preventing slipping, however
useful they would be, as I hold, as early organs of touch. Between
these extremes the _slow loris_ affords a valuable example to study,
with the help of Fig. 61. The foot, as more concerned with prevention
of slipping than the hand, is chosen for observation, but with little
exception the hand agrees closely with it. On the tips of four digits,
D 1, 3, 4 and 5, omitting D 2 for the moment, the ridges are arranged
nearly in a longitudinal direction, and would on that account have
little or no effect in preventing slipping of the foot. If this be
disputed one can but reply that if the need of preventing slipping
in this tiny area were to call forth selective value this is not the
arrangement of the ridges that best serves the purpose. It may be
remarked here that the pulps of _lemurs_, the _marmoset_ and _squirrel
monkey_ all show this indifferent mode of grouping of ridges. The
aborted D 2 of the _loris_, with its hooked nail overhanging the
circular pattern of ridges, is obviously quite unadapted for any
non-slipping effect of its skin, as a glance at the figure shows. On
the remaining segments of the digits the ridges in the main slope from
each side of each digit in the distal direction and fail here also to
obtain the best, or transverse direction for preventing slipping in
locomotion. The corresponding surface of D 1 is not different from its
pulp as to direction of ridges, and it is here to be noted and admitted
that when this muscular great toe is tightly applied to a branch,
which from its shape it must cross at a right angle, the non-slipping
effect of the longitudinal ridges would be very effective. One must
then notice that over the middle of the sole of this foot the ridges
have again changed their direction and lie in a transverse direction.
Between this and the basis of the digits are three fleshy pads and an
intervening area of longitudinal ridges.
Public-domain text, read in full here on John Shaqi.
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