Weismann’s work on the germ-plasm in pursuance of a theory of heredity
is pronounced by Romanes to have remained up to 1893 substantially
unaltered, though largely added to in matters of detail, and at the
present time as far as I gather from a study of the more recent
literature this theory holds the field or at least a commanding
position in it.[26] Originally he held that the germ-plasm possessed
_perpetual_ continuity since the first origin of life, and _absolute_
stability since the first origin of sexual propagation, but he has
shown himself willing to surrender the first postulate, and has himself
altered the second. As it stands now it must be admitted that the
continuity of the germ-plasm is an interrupted continuity with the
appearance of every inherited change; the continuity is theoretical,
not actual, and the stability of the germ-plasm is not absolute but of
a high degree. We can thus see in the story of this original theory of
heredity the lighthouse value of the _pharos_ of Ptolemy II.
[26] Romanes, _Examination of Weismannism_, p. 115.
“It is doubtful if anything better as to Weismann’s theory of
heredity can be said to-day than Romanes said in 1893, and inasmuch
as these two latter or distinctive postulates are not needed for
Weismann’s theory of heredity, while they are both essential to his
theory of evolution, I cannot but regret that he should have thus
crippled the former by burdening it with the latter. Hence my object
throughout has been to display, as sharply as possible, the contrast
that is presented between the brass (“iron” preferably) and the clay
in the colossal figure which Weismann has constructed. Hence also
my emphatic dissent from his theory of evolution does not prevent
me from sincerely appreciating the great value which attaches to
his theory of heredity. And although I have not hesitated to say
that this theory is, in my opinion, incomplete; that it presents not
a few manifest inconsistencies, and even logical contradictions;
that the facts on which it is founded have always been facts of
general knowledge; that in all its main features it was present to
the mind of Darwin, and distinctly formulated by Galton; that in so
far as it has been constituted the basis of a more general theory
of organic evolution it has proved a failure; such considerations
in no way diminish my cordial recognition of the services which its
distinguished author has rendered to science by his speculations
upon these topics. For not only has he been successful in drawing
renewed and much more general attention to the important questions
touching the transmissibility of acquired characters, the causes of
variation, and so on; but even those parts of his system which have
proved untenable are not without such value as temporary scaffoldings
present in relation to permanent buildings. Therefore, if I have
appeared to play the _role_ of a hostile critic, this has been only
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