As just stated, the filaments are not glandular, and do not secrete.
Nor have they the power of absorption, as may be inferred from drops of
a solution of carbonate of ammonia (one part to 146 of water), placed
on two filaments, not producing any effect on the contents of their
cells, nor causing the lobes to close, When, however, a small portion
of a leaf with an attached filament was cut off and immersed in the
same solution, the fluid within the basal cells became almost instantly
aggregated into purplish or colourless, irregularly shaped masses of
matter. The process of aggregation gradually travelled up the filaments
from cell to cell to their extremities, that is in a reverse course to
what occurs in the tentacles of Drosera when their glands have been
excited. Several other filaments were cut off close to their bases, and
left for 1 hr. 30 m. in a weaker solution of one part of the carbonate
to 218 of water, and this caused aggregation in all the cells,
commencing as before at the bases of the filaments.
Long immersion of the filaments in distilled water likewise causes
aggregation. Nor is it rare to find the contents of a few of the
terminal cells in a spontaneously aggregated condition. The aggregated
[page 291] masses undergo incessant slow changes of form, uniting and
again separating; and some of them apparently revolve round their own
axes. A current of colourless granular protoplasm could also be seen
travelling round the walls of the cells. This current ceases to be
visible as soon as the contents are well aggregated; but it probably
still continues, though no longer visible, owing to all the granules in
the flowing layer having become united with the central masses. In all
these respects the filaments of Dionaea behave exactly like the
tentacles of Drosera.
Notwithstanding this similarity there is one remarkable difference. The
tentacles of Drosera, after their glands have been repeatedly touched,
or a particle of any kind has been placed on them, become inflected and
strongly aggregated. No such effect is produced by touching the
filaments of Dionaea; I compared, after an hour or two, some which had
been touched and some which had not, and others after twenty-five
hours, and there was no difference in the contents of the cells. The
leaves were kept open all the time by clips; so that the filaments were
not pressed against the opposite lobe.
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