the species of Droseraceae absorb fluids or at least allow them readily
to enter,* it seemed desirable to ascertain how far the glands of other
plants which are not specially adapted for capturing insects, had the
same power. Plants were chosen for trial at hazard, with the exception
of two species of saxifrage, which were selected from belonging to a
family allied to the Droseraceae. Most of the experiments were made by
immersing the glands either in an infusion of raw meat or more commonly
in a solution of carbonate of ammonia, as this latter substance acts so
powerfully and rapidly on protoplasm. It seemed also particularly
desirable to ascertain whether ammonia was absorbed, as a small amount
is contained in rain-water. With the Droseraceae the secretion of a
viscid fluid by the glands does not prevent their absorbing; so that
the glands of other plants might excrete superfluous matter, or secrete
an odoriferous fluid as a protection against the attacks of insects, or
for any other purpose, and yet have the power of absorbing. I regret
that in the following cases I did not try whether the secretion could
digest or render soluble animal substances, but such experiments would
have been difficult on account of the small size of the glands and the
small amount of secretion. We shall see in the next chapter that the
secretion from the glandular hairs of Pinguicula certainly dissolves
animal matter.
[Saxifraga umbrosa.—The flower-peduncles and petioles of the leaves are
clothed with short hairs, bearing pink-coloured glands, formed of
several polygonal cells, with their pedicels divided by partitions into
distinct cells, which are generally colourless, but sometimes pink. The
glands secrete a yellowish viscid fluid, by
* The distinction between true absorption and mere permeation, or
imbibition, is by no means clearly understood: see Müller’s
‘Physiology,’ Eng. translat. 1838, vol. i. p. 280. [page 346]
which minute Diptera are sometimes, though not often, caught.* The
cells of the glands contain bright pink fluid, charged with granules or
with globular masses of pinkish pulpy matter. This matter must be
protoplasm, for it is seen to undergo slow but incessant changes of
form if a gland be placed in a drop of water and examined. Similar
movements were observed after glands had been immersed in water for 1,
3, 5, 18, and 27 hrs. Even after this latter period the glands retained
their bright pink colour; and the protoplasm within their cells did not
appear to have become more aggregated. The continually changing forms
of the little masses of protoplasm are not due to the absorption of
water, as they were seen in glands kept dry.
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