As we have already seen reason to believe, in the course of organic development when the limits of size and efficiency in the unicellular organism were reached, the only possible access of advantage to the competing organism was gained by the appearance of combination. In the scale of the metazoa {61} we see the advantages of combination and division of labour being more and more made use of, until the individual cells lose completely the power of separate existence, and their functions come to be useful only in the most indirect way and through the organisms of which the cells are constituents. This complete submergence of the cell in the organism indicates the attainment of the maximum advantages to be obtained from this particular access in complexity, and it indicates to us the direction in which development must proceed within the limits which are produced by that other access of complexity—gregariousness. The success and extent of such development clearly depend on the relation of two series of activities in the individual which may in the most general way be described as the capacity for varied reaction and the capacity for communication. The process going on in the satisfactorily developing gregarious animal is the moulding of the varied reactions of the individual into functions beneficial to him only indirectly through the welfare of the new unit—the herd. This moulding process is a consequence of the power of intercommunication amongst the individual constituents of the new unit. Intercommunication is thus seen to be of cardinal importance to the gregarious, just as was the nervous system to the multicellular. Moreover, in a given gregarious species the existence of a highly developed power of reaction in the individual with a proportionately less developed capacity for communication will mean that the species is not deriving the advantages it might from the possession of gregariousness, while the full advantages of the type will be attained only when the two sets of activities are correspondingly strong. Here we may see perhaps the explanation of the astounding success and completeness of {62} gregariousness in bees and ants. Their cycle of development was early complete because the possibilities of reaction of the individual were so small, and consequently the capacity for intercommunication of the individual was relatively soon able to attain a corresponding grade. The individual has become as completely merged in the hive as the single cell in the multicellular animal, and consequently the whole of her activities is available for the uses of the State. It is interesting to notice that, considered from this aspect, the wonderful society of the bee, with its perfect organization and its wonderful adaptability and elasticity, owes its early attainment of success to the smallness of the brain power of the individual.