But radicles of seedlings, as well as those of more vigorous plants,
would pass over stones, roots and other obstacles, which they must
necessarily encounter in the soil. This they are abundantly able to do,
for they are exceedingly sensitive just above their apices, and bend
like a tendril _towards_ the touching object. When, however, one side
of the apex is pressed by any object, the growing part bends _away_
from that object, and this seems a beautiful adaptation for avoiding
obstacles in the soil, and for following the lines of least resistance.
[Illustration: SEEDLING OF WINTER GRAPE.
Earth Cut Away to Show Directions Taken by Tip of Radicle in Avoiding a
Stone.]
So feeble is the circumnutating movement of the terminal growing part,
both of the primary and secondary radicles, that it can assist them
but little in penetrating the ground, excepting when the superficial
layer is very soft and moist. But it must aid them materially when
they chance to break obliquely into cracks, or into burrows that
have been made by earth-worms or larvæ. Moreover, combined as it is
with the sensitiveness of the tip of the radicle to contact, it can
hardly fail to be of the highest importance, for as the tip is always
endeavoring to bend to all sides, it will press on all sides, and will
thus be able to discriminate between the harder and softer adjoining
surfaces. Consequently, it will tend to bend from the harder soil,
and will thus take the directions of the least resistance. So it will
act if it meet with a stone or the root of another plant in the soil,
as must incessantly occur. If the tip were not sensitive, and did not
excite the upper part of the radicle to bend away, whenever obstacles
were encountered at right angles to its growing direction, it would
undoubtedly be liable to be doubled up into a contorted mass. But with
radicles growing down inclined plates of glass, as shown by experiment,
it has been observed that as soon as the tip merely touched a slip
of wood cemented across the plate, the entire terminal growing point
curved away, so that the tip soon stood at right angles to its former
direction; and thus, as far as the pressure of the surrounding soil
would permit, would it be with an obstacle encountered in the ground.
Thick and strong radicles, like those of the horse-chestnut, are
endowed with less sensitiveness than more delicate ones, and would
therefore be the better able by the force of their growth to overcome
any slight impediment to their progress. Further, as radicles perceive
an excess of moisture in the air on one side and bend towards this
side, it is reasonable to infer that they will act in a similar
manner with respect to moisture in the earth, for the sensitiveness
of moisture resides in the tip, which determines the bending of the
upper part. May not this capacity partly account for the extent to
which drain-pipes often become choked with roots? The direction which
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