Introduction to the Study of Palæontological BotanyBalfour, John Hutton
Science
Introduction to the Study of Palæontological Botany
Balfour, John Hutton
Paleobotany
The part of the bark immediately below the epiphlœum is well
developed, and is of a spongy consistence. When examined
microscopically it is seen to be composed of cells of various
shapes--some elongated fusiform, others rhomboidal, others with
pointed appendages. The variety of forms is very great, but it is
possible that this may be partly owing to the effects of frost on the
cells. On the spontaneous separation of the bark, the portion below
the epiphlœum was seen to consist of distinct plates of a more or
less quadrilateral form, with some of the edges concave and others
convex, a part in the centre indicating the connection with the leaf,
along with which it is detached. In Fig. 6 a leaf is shown with one
of these plates attached.
[Illustration: Fig. 6.]
[Sidenote: Fig. 6. Leaf of Araucaria with a portion of bark.]
The appearances presented by the outer and middle bark of Araucaria
imbricata bear a marked resemblance to those exhibited by certain
fossils included in the genera Sigillaria and Lepidodendron. The
sculpturesque markings on the stems of these fossil plants indicate
their alliance to the ferns and lycopods of the present epoch. But
it is evident, from these markings, that much caution is required
in making this determination. Other points of structure must be
examined before a proper decision can be formed. When, for instance,
the presence of scalariform tissue, or of punctated woody tissue,
has been satisfactorily shown under the microscope, we are entitled
to hazard an opinion as to the affinities of the fossils. In many
instances, however, external appearances are the only data on which
to rely for the determination of fossil genera and species; and rash
conclusions have often been drawn by geologists who have not been
conversant with the structure of plants. The Araucaria markings point
out the need of care in drawing conclusions, and their variations at
different parts of the bark indicate the danger of a rash decision
as to species. There can be no doubt that in vegetable Palæontology
the number of species has been needlessly multiplied--any slight
variation in form having been reckoned sufficient for specific
distinction. We can conceive that the Araucaria bark markings in a
fossil state might easily supply several species of Lepidodendron.
A naturalist, with little knowledge of the present flora of the
globe, ventures sometimes to decide on an isolated fragment. Hence
the crude descriptions of fossil vegetable forms, and the confusion
in which Palæophytology is involved. Every geologist who examines
fossil plants ought to be well acquainted with the minute structure
of living plants, the forms of their roots, stems, leaves, fronds,
and fructification; the markings on the outer and inner surfaces of
their barks, on their stems, and on their rhizomes; the localities in
which they grow, and the climates which genera and species affect in
various parts of the world. (Professor Balfour in the Proceedings of
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