Introduction to the Study of Palæontological Botany — John Shaqi
Introduction to the Study of Palæontological BotanyBalfour, John Hutton
Science
Introduction to the Study of Palæontological Botany
Balfour, John Hutton
Paleobotany
In the present day, acrogenous plants are represented by cellular
and vascular Cryptogams. In considering fossil plants our attention
is specially directed to the latter. In the recent Floras, vascular
Acrogens are represented by such plants as Ferns, Lycopods, and
Equisetums. Some of them have an arborescent habit, but the greater
number are shrubby and herbaceous. Many of them have creeping
rhizomes, which are either subterranean, or run along the surface
of the ground. One of these arborescent forms is seen in Tree-ferns
(Fig. 9). Another form with a rhizome is seen in Fig. 10. The trunks
of ferns are marked by scars, which indicate the parts where the
bases of the fronds were attached, and where the vascular tissue
passes out from the interior (Fig. 11, _a_ and _b_). A transverse
section of the stem (Fig. 12) shows a continuous cylinder of
scalariform vessels (Fig. 13), enclosing a large mass of cellular
tissue frequently penetrated by small scalariform bundles. The
cylinder is pierced by meshes, from the inner sides of which rise the
vascular bundles going to the leaves, while some of the free bundles
of the axis pass through the mesh, carrying with them a portion of
the cellular tissue into the petiole. The fructification consists
of spore-cases (sporangia), often with an elastic ring round them,
containing spores in their interior (Fig. 14).
[Illustration: Fig. 9.]
[Sidenote: Fig. 9. Tree-fern, with a slender cylindrical trunk and a
crown of drooping fronds. It is a vascular acrogen.]
[Illustration: Fig. 10.]
[Sidenote: Fig. 10. _Asplenium_; a species of Spleenwort. A. Rhizome,
_r_, covered with the bases (stalks or stipes) of the fronds; _f_,
fronds in bud, rolled up in a circinate manner (this is very rarely
seen in fossil ferns); _g_, fronds bearing fructification on their
backs. B. Portion of a frond separated to show the linear sori or
clusters of sporangia (spore-cases).]
Among Acrogens of the present day there are also plants belonging
to the natural order Lycopodiaceæ or Club-mosses (Fig. 15), having
creeping stems, which give rise to leafy branches. The leaves are
small, sessile, and moss-like, and the fructification consists of two
kinds of cellular bodies, small spores or microspores (Fig. 16),
and large spores or macrospores (Fig. 17). They consist of cellular
and vascular tissues, the latter occurring in the form of woody,
annular, and scalariform vessels, which occupy the axis or central
part of the stem. They differ from ferns in the distribution of
their vascular bundles. The order is represented also by such plants
as Selaginella, Psilotum, Phylloglossum, and Isoetes. In the plant
called Isoetes (Quillwort) there is a peculiar short stem which does
not increase in height. It produces additions laterally, so that the
stem increases in thickness. The leaves continue to multiply, and
bear fructification at their bases. They have both large and small
spores.
[Illustration: Fig. 11, _a_. Fig. 11, _b_. Fig. 12.]
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