Introduction to the Study of Palæontological BotanyBalfour, John Hutton
Science
Introduction to the Study of Palæontological Botany
Balfour, John Hutton
Paleobotany
Fossil plants are by no means so easily examined as recent species.
They are seldom found in a complete state. Fragments of stems,
leaves, and fruits, are the data by which the plant is to be
determined. It is very rare to find any traces of the flowers. The
parts of fossil plants are usually separated from each other, and
it is difficult to ascertain what are the portions which should be
associated together so as to complete an individual plant. Specimens
are sometimes preserved, so that the anatomical structure of the
organs, especially of the stem, can be detected by very thin slices
placed under the microscope. In the case of some stems the presence
of punctated woody tissue (Fig. 1) has proved of great service as
regards fossil Botany; this structure, along with the absence of
large pitted ducts, serving to distinguish Conifers. The presence
of scalariform vessels indicates a plant belonging to the vascular
Cryptogams, of which the fern is the best known example. The cautions
to be observed in determining fossil plants are noticed by Dr. Hooker
in the Memoirs of the Geological Survey of Great Britain (vol. ii.
p. 387). At the present day, the same fern may have different forms
of fronds, which, unless they were found united, might be reckoned
distinct genera; and remarkable examples are seen in Niphobolus
rupestris and Lindsæa cordata. Moreover, we find the same form of
frond belonging to several different genera, which can only be
distinguished by the fructification; and as this is rarely seen in
fossil ferns, it is often impossible to come to a decided conclusion
in regard to them. A leaf of Stangeria paradoxa was considered
by an eminent botanist as a barren fern frond, but it ultimately
proved to be the leaf of a Cycad. The leaf of Cupania filicifolia,
a Dicotyledon, might easily be mistaken for that of a fern; it
resembles much the frond of a fossil fern called Coniopteris. The
diverse leaves of Sterculia diversifolia, if seen separately, might
easily be referred to different plants. In the same fern we meet
also with different kinds of venation in the fronds. Similar remarks
may be made in regard to other plants. Harvey has pointed out many
difficulties in regard to sea-weeds.
As regards the materials for a fossil flora, the following remarks of
Hugh Miller deserve attention:--
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