Irritability : $b a physiological analysis of the general effect of stimuli in living substanceVerworn, Max
Science
Irritability : $b a physiological analysis of the general effect of stimuli in living substance
Verworn, Max
Irritability
which is dependent upon the intensity of the stimulus._
We have to the present considered only the _simplest_ conditions
existing as a result of the effect of a _single_ series of stimuli and
also of the interference of its individual members. These elementary
conditions are at the basis of an understanding of complicated
_interference effects which arise when two series of stimuli interact_.
In that these processes can be readily explained by the elementary
processes previously described, I will, therefore, dwell but briefly
on this subject. From the standpoint already taken it may be readily
presumed what will happen when two series of stimuli act upon the same
system.
When there is interference of _two series of stimuli_, there are
two resultant possibilities. In one type the stimuli of the one are
active simultaneously with that of the other. In this instance both
stimuli would act as a single stimulus of greater intensity, and we
have essentially the same condition as exists when a single series is
operative. Nevertheless, such cases are practically hardly realized
in the physiological happenings of the organism. More often a state
exists wherein the single stimuli of one series occur in the intervals
of the stimuli of the other. In these cases there is an increase in
the frequency of the stimuli applied in a given length of time. We
have here, then, in principle the same conditions as when a series
of greater frequency is operative. (Figure 55.) The effect of such
alteration in the frequency consists in an increase of the velocity
of the development of summation or inhibition, as the general scheme
(Figure 55) has shown us. Depending upon the special combination
of the factors involved in interference, we may have a summation
of the exciting effect of each series of stimuli or an inhibition
of one series by the exciting effects of the other series. If the
frequency of both series is essentially different, we may have here
the conditions for periodically increasing and decreasing excitations.
Nevertheless these conditions have not been systematically analyzed and
experimentally studied.
[Illustration: A B
Fig. 55.
Interference of two series of stimuli. A--Effect of the one series
alone. Development of tonus by summation. The dots below the curve
indicate the points of time at which the stimuli of the second series
will operate. B--Effect resulting from the interference of both series.
By the addition of the second series the frequency has been doubled.
The result consists in an inhibition. ]
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