Island Life; Or, The Phenomena and Causes of Insular Faunas and FlorasWallace, Alfred Russel
Science
Island Life; Or, The Phenomena and Causes of Insular Faunas and Floras
Wallace, Alfred Russel
Biogeography; Glacial epoch; Island ecology
Overlapping areas among the species of a genus is a more common phenomenon,
and is almost universal where these species are numerous in the same
continent. It is, however, exceedingly irregular, so that we often find one
{19} species extending over a considerable portion of the area occupied by
the genus and including the entire areas of some of the other species. So
little has been done to work out accurately the limits of species that it
is very difficult to give examples. One of the best is to be found in the
genus _Dendroeca_, a group of American wood-warblers. These little birds
all migrate in the winter into the tropical regions, but in the summer they
come north, each having its particular range. Thus, _D. dominica_ comes as
far as the middle Eastern States, _D. coerulea_ keeps west of the
Alleghanies, _D. discolor_ comes to Michigan and New England; four other
species go farther north in Canada, while several extend to the borders of
the Arctic zone.
_The Species of Tits as Illustrating Areas of Distribution._--In our own
hemisphere the overlapping of allied species may be well illustrated by the
various kinds of titmice, constituting the genus Parus, several of which
are among our best known English birds. The great titmouse (_Parus major_)
has the widest range of all, extending from the Arctic circle to Algeria,
Palestine, and Persia, and from Ireland right across Siberia to the Ochotsk
sea, probably following the great northern forest belt. It does not extend
into China and Japan, where distinct species are found. Next in extent of
range is the coal tit (_Parus ater_) which inhabits all Europe from the
Mediterranean to about 64deg N. latitude, in Asia Minor to the Lebanon and
Caucasus, and across Siberia to Amoorland and Japan. The marsh tit (_Parus
palustris_) inhabits temperate and south Europe from 61deg N. latitude in
Norway to Poland and South-west Russia, and in the south from Spain to Asia
Minor. Closely allied to this--of which it is probably only a variety or
sub-species--is the northern marsh tit (_Parus borealis_), which overlaps
the last in Norway and Sweden, and also in South Russia and the Alps, but
extends further north into Lapland and North Russia, and thence probably in
a south-easterly direction across Central Asia to North China. Yet another
closely-allied species (_Parus camtschatkensis_) ranges from North-eastern
Russia across Northern Siberia to Lake Baikal and to Hakodadi in Japan,
thus overlapping _Parus borealis_ in the {20} western portion of its area.
Our little favourite, the blue tit (_Parus coeruleus_) ranges over all
Europe from the Arctic circle to the Mediterranean, and on to Asia Minor
and Persia, but does not seem to pass beyond the Ural mountains. Its lovely
eastern ally the azure tit (_Parus cyaneus_) overlaps the range of _P.
coeruleus_ in Western Europe as far as St. Petersburg and Austria, rarely
straggling to Denmark, while it stretches all across Central Asia between
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