Island Life; Or, The Phenomena and Causes of Insular Faunas and FlorasWallace, Alfred Russel
Science
Island Life; Or, The Phenomena and Causes of Insular Faunas and Floras
Wallace, Alfred Russel
Biogeography; Glacial epoch; Island ecology
But here again we meet with exactly the same difficulty as in the mammalia,
in the comparative poverty of the types of birds now inhabiting Celebes.
Although the preponderance of affinity, especially in the case of its more
ancient and peculiar forms, is undoubtedly with Asia rather than with
Australia; yet, still more decidedly than in the case of the mammalia, are
we forbidden to suppose that it ever formed a part of the old Asiatic
continent, on account of the _total_ absence of so many important and
extensive groups of Asiatic birds. It is not single species or even genera,
but whole families that are thus absent, and among them families which are
pre-eminently characteristic of all tropical Asia. Such are the Timaliidae,
or babblers, of which there are twelve genera in Borneo, and nearly thirty
genera in the Oriental Region, but of which one species only, hardly
distinguishable from a Malayan form, inhabits Celebes; the Phyllornithidae,
or green bulbuls, and the Pycnonotidae, or bulbuls, both absolutely
ubiquitous in tropical Asia and Malaya, but unknown in Celebes; the
Eurylaemidae, or gapers, found everywhere in the great Malay Islands; the
Megalaemidae, or barbets; the Trogonidae, or trogons; and the Phasianidae,
or pheasants, all pre-eminently Asiatic and Malayan but all absent from
Celebes, with the exception of the common jungle-fowl, which, owing to the
passion of Malays for cock-fighting, may have been introduced. To these
important _families_ may be added Asiatic and Malayan _genera_ by the
score; but, confining ourselves to these seven ubiquitous families, we must
ask,--Is it possible, that, at the period when the ancestors of the
peculiar Celebes mammals entered the island, and when the forms of life,
though distinct, could not have been quite unlike those now living, it
could have actually formed a part of the continent without {461} possessing
representatives of the greater part of these extensive and important
families of birds? To get rid altogether of such varied and dominant types
of bird-life by any subsequent process of submersion is more difficult than
to exterminate mammalia; and we are therefore again driven to our former
conclusion--that the present land of Celebes has never (in Tertiary times)
been united to the Asiatic continent, but has received its population of
Asiatic forms by migration across narrow straits and intervening islands.
Taking into consideration the amount of affinity on the one hand, and the
isolation on the other, of the Celebesian fauna, we may probably place the
period of this earlier migration in the early part of the latter half of
the Tertiary period, that is, in middle or late Miocene times.
Public-domain text, read in full here on John Shaqi.
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