Island Life; Or, The Phenomena and Causes of Insular Faunas and FlorasWallace, Alfred Russel
Science
Island Life; Or, The Phenomena and Causes of Insular Faunas and Floras
Wallace, Alfred Russel
Biogeography; Glacial epoch; Island ecology
_Celebes not Strictly a Continental Island._--A study of the mammalian and
of the bird-fauna of Celebes thus leads us in both cases to the same
conclusion, and forbids us to rank it as a strictly continental island on
the Asiatic side. But facts of a very similar character are equally opposed
to the idea of a former land-connection with Australia or New Guinea, or
even with the Moluccas. The numerous marsupials of those countries are all
wanting in Celebes, except the phalangers of the genus Cuscus, and these
arboreal creatures are very liable to be carried across narrow seas on
trees uprooted by earthquakes or floods. The terrestrial cassowaries are
equally absent; and thus we can account for the presence of all the
Moluccan or Australian types actually found in Celebes without supposing
any land-connection on this side during the Tertiary period. The presence
of the Celebes ape in the island of Batchian, and of the babirusa in Bouru,
can be sufficiently explained by a somewhat closer approximation of the
respective lands, or by a few intervening islands which have since
disappeared, or it may even be due to human agency.
If the explanation now given of the peculiar features presented by the
fauna of Celebes be the correct one, we are fully justified in classing it
as an "anomalous island," since it possesses a small but very remarkable
mammalian fauna, without ever having been directly united with any {462}
continent or extensive land; and, both by what it has and what it wants,
occupies such an exactly intermediate position between the Oriental and
Australian regions that it will perhaps ever remain a mere matter of
opinion with which it should properly be associated. Forming, as it does,
the western limit of such typical Australian groups as the Marsupials among
mammalia, and the Trichoglossidae and Meliphagidae among birds, and being
so strikingly deficient in all the more characteristic Oriental families
and genera of both classes, I have always placed it in the Australian
Region; but it may perhaps with equal propriety be left out of both till a
further knowledge of its geology enables us to determine its early history
with more precision.
_Peculiarities of the Insects of Celebes._--The only other class of animals
in Celebes, of which we have a tolerable knowledge, is that of insects,
among which we meet with peculiarities of a very remarkable kind, and such
as are found in no other island on the globe. Having already given a full
account of some of these peculiarities in a paper read before the Linnean
Society--republished in my _Contributions to the Theory of Natural
Selection_,--while others have been discussed in my _Geographical
Distribution of Animals_ (Vol. I. p. 434)--I will only here briefly refer
to them in order to see whether they accord with, or receive any
explanation from, the somewhat novel view of the past history of the island
here advanced.
Public-domain text, read in full here on John Shaqi.
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