Island Life; Or, The Phenomena and Causes of Insular Faunas and FlorasWallace, Alfred Russel
Science
Island Life; Or, The Phenomena and Causes of Insular Faunas and Floras
Wallace, Alfred Russel
Biogeography; Glacial epoch; Island ecology
Among reptiles we have two families of snakes--the Dendrophidae or
tree-snakes, and the Dryiophidae or green whip-snakes--which are also found
in the three tropical regions of Asia, Africa, and America, but in these
cases even some of the genera are common to Asia and Africa, or to Africa
and America. The lizards forming the family Amphisbaenidae are divided
between tropical Africa and America, a few species only occurring in the
southern portion of the adjacent temperate regions; while even the
peculiarly American family of the iguanas is represented by two genera in
Madagascar, and one in the Fiji and Friendly Islands. Passing on to the
Amphibians the worm-like Caeciliadae are tropicopolitan, as are also the
toads of the family Engystomatidae. Insects also furnish some analogous
cases, three genera of Cicindelidae, (Pogonostoma, Ctenostoma, and
Peridexia) showing a decided connection between this family in South
America and Madagascar; while the beautiful family of diurnal moths,
Uraniidae, is confined to the same two countries. A somewhat similar but
better known illustration is afforded by the two genera of ostriches, one
confined to Africa and Arabia, the other to the plains of temperate South
America.
_General features of Overlapping and Discontinuous Areas._--These numerous
examples of discontinuous genera and families form an important section of
the facts of animal dispersal which any true theory must satisfactorily
account for. In greater or less prominence they are to be found all over
the world, and in every group of animals, and they grade imperceptibly into
those cases of conterminous and overlapping areas which we have seen to
{29} prevail in most extensive groups of species, and which are perhaps
even more common in those large families which consist of many closely
allied genera. A sufficient proof of the overlapping of generic areas is
the occurrence of a number of genera of the same family together. Thus in
France or Italy about twenty genera of warblers (Sylviadae) are found, and
as each of the thirty-three genera of this family inhabiting temperate
Europe and Asia has a different area, a great number must here overlap. So,
in most parts of Africa, at least ten or twelve genera of antelopes may be
found, and in South America a large proportion of the genera of monkeys of
the family Cebidae occur in many districts; and still more is this the case
with the larger bird families, such as the tanagers, the tyrant shrikes, or
the tree-creepers, so that there is in all these extensive families no
genus whose area does not overlap that of many others. Then among the
moderately extensive families we find a few instances of one or two genera
isolated from the rest, as the spectacled bear, Tremarctos, found only in
Chili, while the remainder of the family extends from Europe and Asia over
North America to the Mountains of Mexico, but no further south; the
Bovidae, or hollow-horned ruminants, which have a few isolated genera in
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